生态位因子分析在大熊猫(Ailuropoda melanoleuca)生境评价中的应用

生态位因子分析是研究物种地理分布的一种多变量分析方法,其最大优点是模型计算只需物种"出现点"的数据,而不需要"非出现点"数据,在生境评价与生境预测中得到广泛应用.将该方法应用于大熊猫生境适宜性评价中,利用大熊猫活动痕迹点和遥感数据分析了平武县大熊猫生境分布现状,综合评价了该县自然保护区的分布状况和存在的保护空缺.研究结果表明,大熊猫偏好在中高海拔(＞2128 m)的针叶林和针阔混交林中活动,而避免在落叶阔叶林和灌丛林中活动,避免在有人为干扰的地区活动,农田是对大熊猫活动影响强度最大的人为干扰因子.平武县大熊猫生境主要分布在该县西部和北部地区,总面积为234033 hm2,其中适宜生境为106345 hm2,次适宜生境为127688 hm2.目前该县已建的3个大熊猫自然保护区使47.2%的大熊猫生境得到保护(包括49.2%的适宜生境和45.6%的次适宜生境),尽管如此,保护区之间连接性差,存在严重的保护空缺.该县北部的白马乡、木座乡是大熊猫的主要分布区,却没有得到保护区的有效保护,建议在该地区新建自然保护区.     

蜂桶寨自然保护区大熊猫生境适宜性评价与保护管理对策

对濒危野生动物而言,开展生境质量评价是完善就地保护措施、提高自然保护成效的基础。蜂桶寨自然保护区位于邛崃山系中段,是大熊猫模式标本的采集地。而此前有关该保护区大熊猫生境的研究主要集中在生境的选择上,生境适宜性评价方面的研究尚未开展。本文以海拔、坡度、植被类型、食物来源等4个生态因子为指标,通过构建景观连接度模型对保护区内大熊猫生境适宜性进行评价。结果表明,保护区内大熊猫不适宜生境面积、较适宜生境面积及最适宜生境面积分别为223.26、133.67和33.46 km2,分别占保护区总面积的57%、34%和9%。从空间分布来看,最适宜生境和较适宜生境在保护区内的分布较为破碎分散,大熊猫活动痕迹点更多地集中在生境适宜程度较高的区域。为加强该区大熊猫种群的科学保护,应控制敏感区域的人为活动,确保保护区内大熊猫个体在不同生境斑块之间迁移扩散。日常保护管理工作中应重点关注与黄水河自然保护区相连区域,并可适度扩大蜂桶寨自然保护区的管护范围。     

青木川自然保护区大熊猫生境评价

在实地调查和收集资料的基础上,对陕西省青木川自然保护区大熊猫生境选择的特点进行研究;利用层次分析法以及ArcMap、ArcView等地理信息系统辅助软件,从地理环境因素、生物群落因素、主食竹因素和干扰因素4方面对大熊猫栖息地的核心区和缓冲区进行质量评价.结果表明：研究区适宜大熊猫生存的生境面积比例为62.1%;次适宜生境面积占31.0%;不适宜生境面积占6.9%.大熊猫适宜生境主要分布在研究区西部和北部海拔较高的地带,不适宜生境主要受植被类型、竹子因素、干扰因素等影响.为减少大熊猫不适宜生境,需要保护区管理者进一步完善管理机制,控制人类活动,特别是海拔900～1600 m范围内的放牧、割竹和砍柴活动.     

甘肃大熊猫生境质量评价

本研究以第四次大熊猫(Ailuropoda melanoleuca)调查、甘肃省森林资源二类清查等数据为基础资料,依据国内外有关大熊猫对生境选择利用的研究成果,结合本地本山系大熊猫分布及活动的实际情况,选取竹子分布、植被类型、海拔、坡度、坡向、道路和居民区等评价因子,建立了大熊猫生境质量评价指标体系和准则,借助Arcgis10.2软件,利用栅格赋值和空间叠加原理系统地研究了甘肃大熊猫生境质量。结果表明,甘肃大熊猫生境质量总体较好,适宜和较适宜生境面积总和占到了69.77%;四个局域种群单元中白水江单元适宜生境面积最大,其次为甘南单元;各大熊猫分布县中,文县的大熊猫生境质量最好,其次为舟曲;甘肃大熊猫适宜生境大都处于保护区中,各保护区适宜生境总和为178 771 hm~2,占全省大熊猫适宜生境的88.72%;各保护区中,白水江国家级自然保护区的适宜生境面积最大,其次为插岗梁省级保护区。     

大相岭山系大熊猫生境评价与保护对策研究

从长远来看,从山系或更大尺度上来开展大熊猫(Ailuropodamelanoleuca)生境的研究与保护更有意义。本研究通过野外调查,在地理信息系统(GIS)和遥感(RS)技术支持下,利用大熊猫生境结构理论模型,选取海拔、坡度、植被类型、竹子分布、道路和居民点的分布等评价因子,系统地研究了大相岭山系大熊猫生境的分布、生境质量与空间格局,以及生境保护现状,在此基础上提出了该山系生境保护与自然保护区建设的对策。遥感数据分析结果表明,与大熊猫生境密切相关的阔叶林与针叶林的面积为344,970hm2,占该地区总面积的58.4%,灌丛占18.4%,其他的土地利用与土地覆被类型占23.2%。综合评价结果表明,大相岭山系的大熊猫潜在生境为118,749hm2,由于森林及矿产资源的开发、交通以及农业活动等的影响导致生境面积减少为93,115hm2,且尚存的生境被隔离为两个互相独立的生境单元,使荥河与瓦屋山这两个种群的交流受到严重阻碍,而现有的保护区仅保护了生境总面积的28.0%。为了有效地保护该山系的大熊猫,应该注意从三个方面加强对大熊猫生境的规划与保护:(1)扩大自然保护区的保护范围,使大熊猫生境的集中分布区都得到保护;(2)以退耕还林和天然林保护工程为契机,加强被隔离生境的联系,促进大熊猫种群之间的交流;(3)控制海拔1,800–2,700m之间人类活动对大熊猫生境的影响。     

基于MaxEnt模型的广西黑颈长尾雉潜在栖息地预测与保护空缺分析

了解物种生境的空间分布以及和环境因子的关系,对物种的保护有重要意义。黑颈长尾雉Syrmaticus humiae作为在广西壮族自治区分布的鸟类,通过查阅文献与分布名录得到其分布点,以GIS空间技术运用Max Ent模型对区内黑颈长尾雉的生境适应性和其主要影响因子进行分析,模型评价结果为优秀水平。通过划分出的适宜生境和现有保护区进行保护空缺分析。模型预测黑颈长尾雉的生境适宜区主要分布在广西的西北区域,适宜生境总面积为6 771.84 km~2,已建立的保护区覆盖了16.72%的适宜生境,尚有4 985.86 km2的适宜生境在保护区之外。为更有效地实行保护,应对保护空缺区域进行填补。     

农用地时空变化对大熊猫生境的影响:以岷山中部地区的白草河流域为例

岷山中部地区是我国野生大熊猫(Ailuropoda melanoleuca)种群分布密度较高的地区之一,但近年来砍伐森林、种植药材等农业活动较为突出,可能对大熊猫及其生境利用造成不利影响.为了明确该地区农用地的时空变化特征,及其对保护区内外大熊猫生境的影响,我们采用遥感影像解析、GIS空间分析和野外调查相结合的方法在白草河流域进行了本项研究.结果表明.该流域的农用地主要分布在海拔1,700 m以下的地区.从1994-2008年,农用地面积呈现先减少后增加的趋势:1994-2001年间,农用地面积减少,减少区域主要在海拔1,700 m以下的地区;2001-2008年间,农用地面积增加明显,增加区域主要在海拔1,700m以上的高海拔地区和自然保护区周边.农用地扩张直接导致1994-2008年间该地区5,281 hm2(6.46%)的大熊猫生境丧失.就保护区内外而言,保护区外部大熊猫生境的丧失比例达21.53%,而保护区内部的生境变化不大.为了有效保护该地区的大熊猫生境,除了禁止在自然保护区内部进行开发外,建议对保护区外部的大熊猫生境进行保护,禁止在海拔1,700 m以上的高海拔地区进行土地开发.     

新疆塔什库尔干野生动物自然保护区马可波罗盘羊生境适宜性评价

马可波罗盘羊是帕米尔高原的代表性物种,开展生境适宜性评价对于该物种的保护与管理具有重要意义。综合考虑植被、地形等影响马可波罗盘羊生境选择的关键因子,以及道路和牧场等人为干扰因子,借助ArcGIS,构建生境适宜性指数模型,在不同尺度上评价分析马可波罗盘羊的生境适宜性及其季节变化特征。结果表明:15836 km~2保护区内马可波罗盘羊夏季潜在适宜生境面积为2127.19 km~2,冬季为1915.70 km~2。保护区西北部面积3767.73 km~2的马可波罗盘羊实际分布区内,夏季潜在适宜生境面积为1095.48 km~2,冬季为1072.82 km~2,马可波罗盘羊适宜生境集中分布于保护区内该物种实际分布区。受人为干扰,保护区内马可波罗盘羊夏季和冬季实际适宜生境丧失率分别为18.43%和17.78%,实际分布区内适宜生境丧失率分别为33.65%和29.73%,表明实际分布区马可波罗盘羊适宜生境受人类活动影响较大,应予以重点保护。总体而言,影响马可波罗盘羊生境适宜性的关键因素是国道314和放牧。基于上述研究结果,作者提出了4点关于马可波罗盘羊生境保护,以及保护区管理与规划的建议与思考。     

基于MAXENT模型的秦岭山系黑熊潜在生境评价

明确物种生境空间分布格局及其与环境因子的关系,对制定合理有效的保护对策十分重要.本文以黑熊(Ursus thibetanus)为研究对象,以其重要柄息地秦岭山系为研究区域,选取68个黑熊分布点数据和34个环境因子,应用MAXENT模型分析其生境空间分布及主要影响因子,以最大化Kappa值的生境适宜性指数为阈值划分适宜生境,结合已建自然保护区进行保护空缺分析,并通过构建阻力面和最小成本路径分析,开展黑熊生境廊道规划.结果表明:人类干扰和土地利用类型是影响黑熊生境选择的主要生态因子,居民点密度、到荒草地距离、到耕地距离3个因子对黑熊生境选择有重要影响,其综合贡献值分别为21.4%、17.5%和15.9%,到阔叶林距离、到水体距离等因子次之.黑熊的适宜生境主要集中分布在秦岭山系主脊的中西部地区,占整个秦岭山系面积的19.23%.空缺分析表明:已建自然保护区群覆盖了23.49%的适宜生境,但尚有8,480 km2处于保护区之外.为更有效保护秦岭黑熊及其生境,建议建设12条生境廊道,同时结合其他物种进行系统保护规划.     

岷山地区大熊猫(Ailuropoda melanoleuca)生境影响因子连通性及主导因子

岷山地区是大熊猫保护的关键地区。运用图论分析法研究了岷山地区大熊猫生境影响因子,及其相互作用关系。结果表明：在岷山地区,12个影响因子66对组合中,47.0％的影响因子间存在直接的联系,其中89.2％的影响具有增大效应,这说明人类活动对大熊猫生境的影响具有协同增大效应,并占主导地位。研究发现,在影响因子关系集中,存在强连通性的子集K＝{ (TD);(RB);(MI); (HPL);(AP);(AD);(SR);(TH);（FC）},这表明近期进行的旅游景点开发,公路建设,采矿,高压电线走廊建设,以及传统的农业开发, 畜牧业, 薪柴采集等影响因子之间存在着强烈相互作用关系,这些人类活动还会将其他人类活动对大熊猫生境的影响放大,加剧其对大熊猫生境的不利影响,是影响岷山地区大熊猫生境的主导因素。研究还表明如果不能对主导因子进行有效的控制,很难有效地保护岷山地区大熊猫生境。研究表明图论分析法是研究大熊猫生境影响因子的一个有效工具,有助于明确影响熊猫生境的主导和关键因子,为制定有效的大熊猫保护策略提供科学依据。     

动物的生境选择

动物的生境选择颜忠诚陈永林（中国科学院动物研究所,北京１０００８０）ＨａｂｉｔａｔＳｅｌｅｃｔｉｏｎｉｎＡｎｉｍａｌｓ．ＹａｎＺｈｏｎｇｃｈｅｎｇ,ＣｈｅｎＹｏｎｇｌｉｎ（ＩｎｓｔｉｔｕｔｅｏｆＺｏｏｌｏｇｙ,ＡｃａｄｅｍｉａＳｉｎｉｃａ,Ｂｅｉｊｉ...     

Demography and feeding behavior of the kelp crab Taliepus marginatus in subtidal habitats dominated by the kelps Macrocystis pyrifera or Lessonia trabeculata

We studied the population and feeding ecology of the kelp crab Taliepus marginatus in subtidal kelp forests dominated by either of two morphologically different kelp species (Macrocystis pyrifera or Lessonia trabeculata) in northern Chile. The sizes and abundances of T. marginatus differed between the two kelp habitats. Kelp crabs were more abundant in the M. pyrifera forest than in the L. trabeculata forest. Size‐frequency distributions showed that juvenile and immature crabs were more common in the M. pyrifera forest than in the L. trabeculata forest, where reproductive adults predominated. The smaller crabs in the M. pyrifera habitat also consumed a higher proportion of kelp tissues than the larger crabs in the L. trabeculata habitat, which had a higher proportion of animal food in their diet. In both kelp forests, individuals of T. marginatus showed a similar pattern of nocturnal feeding over a 24‐h period, consuming more food at night than during the day. The more complex and dense forests of M. pyrifera appear to present better nursery habitats for juvenile kelp crabs than the more open and less dense forests dominated by L. trabeculata. These results suggest that the role of the two kelp habitats for T. marginatus varies during the life cycle of the kelp crabs, with M. pyrifera tending to have nursery function and L. trabeculata being more suitable as a reproductive habitat.     

Interhabitat migration of juvenile Atlantic salmon in a Newfoundland river system, Canada

Migrations of juvenile Atlantic salmon between habitats (both fluvial and lacustrine) were studied in a Newfoundland river system from May through September. Salmon parr showed poor site fidelity in fluvial habitats, and high rates of migration, especially in spring-early summer. Most habitat shifts were upstream from fluvial habitats to a lake at the head of a small tributary. In September, 10% of the salmon parr caught in the lake ( n =275) had been marked in the main stem of the river or in the tributary stream (total n =641). The abundance of parr in the lake increased in May-June, then declined gradually to September. Most of the lacustrine immigrants were 1+ parr. Mature males were found amongst the autumnal emigrants. Salmon parr in the lake grew larger than those in the river, and lacustrine smolts captured in the lake were larger than those caught at the mouth of the river.     

濒危乔木红豆杉不同更新阶段的生境特征比较

以福建梅花山自然保护区濒危乔木红豆杉(Taxus chinensis(Pilger)Rehd.)为对象,对红豆杉的种子排放、幼苗更新、幼树更新3个更新阶段的生境特征进行研究,并比较生态位的变化,以此评价红豆杉不同更新阶段的生境需求。结果显示:随着个体发育生长,红豆杉更新生境发生了明显转变,种子排放地与幼苗生境空间一致性较高,而它们与幼树生境存在明显的空间不一致性。从生态位角度来看,种子排放地和幼苗生境重叠指数≤0.5的仅有海拔和坡度2个因子,这说明幼苗的空间格局受到鸟类传播的强烈影响,鸟类传播所排放的种子能萌发并生长成幼苗。幼苗和幼树生境重叠指数≤0.5的因子有海拔、坡向、植被类型,说明在大尺度景观因子和庇护植被层面上,植物的更新需求出现了明显的不一致。研究结果表明红豆杉的更新生境存在明显的阶段变化,可能造成植物更新出现建成限制,使幼树阶段成为更新的瓶颈阶段。     

Simple predictions of instream habitat model outputs for fish habitat guilds in large streams

1. In the context of a generalised modification of hydraulic conditions in medium to large streams, modelling the impacts of stream regulation on fish communities in multiple streams is an important challenge for basic and applied freshwater ecology. Conventional instream habitat models such as PHABSIM link a hydraulic model with preference curves for various species to estimate habitat value changes with discharge in stream reaches. Despite world‐wide applications, they have been scarcely used in multiple sites with multiple species. 2. We assigned 21 size classes of European fish species to four habitat guilds (cluster analysis grouping size classes with comparable microhabitat preference curves). Then, we ran a conventional instream habitat model on 28 French stream reaches belonging to the `barbel zone', to estimate habitat values versus discharge curves for the 21 size classes. We summarised the outputs as mean habitat values for guilds, and tested if they were predictable from average characteristics of reaches (discharge, depth, width, particle size). 3. As was obtained elsewhere for populations, habitat values for guilds were strongly related to average, dimensionless characteristics of reaches. The Reynolds number of reaches, equivalent to a discharge per width unit, reflected most of the discharge‐dependent changes in habitat values (within reaches). In particular, habitat values of species preferring bank (respectively midstream) microhabitats decreased (respectively increased) with increasing Reynolds number. The Froude number at median discharge was the major predictor of reach‐dependent but discharge‐independent variations in habitat values. Habitat values of species preferring riffle versus pool or bank microhabitats were higher in reaches with high Froude numbers. These relationships were consistent with existing knowledge on the different species. 4. Such results suggest that the input variables required to estimate habitat values for fish communities can be greatly simplified, as illustrated by a general estimation of the sensitivity of species preferring midstream habitats to discharge changes in any reach. Cost‐efficient alternatives to conventional instream habitat models should facilitate their validation in multiple sites, a point that remains critical in instream habitat modelling of fish communities.     

官山保护区白颈长尾雉栖息地适宜性评价

2009年4月-2009年5月和2009年12月-2010年1月,在江西官山国家级自然保护区对白颈长尾雉(Syrmaticus ellioti) 繁殖期和越冬期的栖息地进行调查。通过实地调查,结合已有研究结果,确定了官山自然保护区白颈长尾雉栖息地适宜性评价因子,即植被类型、海拔、坡度和坡向等4个栖息地因子。基于"被利用生境-可获得生境比较法",以对各评价因子不同梯度的资源选择指数为依据,确定了白颈长尾雉适宜性评价准则。应用ArcGIS 9.3空间分析功能和栖息地片断化指数,对官山保护区白颈长尾雉栖息地进行了适宜性评价。结果表明:在官山自然保护区适宜白颈长尾雉繁殖的栖息地斑块有502个,总面积为3307 hm²,占保护区总面积的28.6%,分离度指数和片断化程度指数分别为0.365和0.912;适宜白颈长尾雉越冬的栖息地斑块有514个,总面积2755 hm²,占保护区总面积的23.8%,分离度指数和片断化程度指数分别为0.443和0.939。官山保护区白颈长尾雉栖息地片断化较严重,片断化现象在白颈长尾雉越冬期加剧。因此,迫切需要采取措施白颈长尾雉现存栖息地进行保护,特别是要高度重视其越冬栖息地的保护。     

基于生境可获得性的完达山地区马鹿(Cervus elaphus xanthopygus)冬季生境选择

于2004～2005年冬季研究了完达山地区马鹿的生境利用和选择。在野外调查的基础上,获取了研究地区马鹿种群水平上的生境利用数据;利用地理信息系统和遥感技术,从区域尺度上测度了各类生境因子的可获得性。通过比较被利用生境与可获得生境,测度了马鹿冬季对各类生境或生态因子的选择。结果表明,马鹿对植被类型、海拔、坡向、NDVI等级、到公路距离表现出非比例利用,对坡度的利用与其可获得性则没有显著性差异。马鹿选择采伐迹地,低郁闭度生境（第2、3NDVI等级）,200～300m海拔区间,南坡,距离公路600～700m的区间。马鹿采食生境和卧息生境对各类因子具有不同的选择性,表现出不同功能生境分离的特征。对于采食生境,马鹿选择采伐迹地、阔叶疏林,低郁闭度（第2、3NDVI等级）,南坡;对于卧息生境,马鹿仅选择采伐迹地,低郁闭度（第2、3、4NDVI等级）。根据马鹿活动样点的出现与否,建立了马鹿冬季生境选择的逻辑斯谛回归模型。模型回判结果表明,该模型对马鹿活动样方与对照样方的总体正确预测率为74．4％,对马鹿活动样方的正确预测率为84．2％,对对照样方的正确预测率为62．4％,能够较好的预测生境利用概率。     

Habitat characteristics of wintering Wood Warbler Phylloscopus sibilatrix in the Centre Region of Cameroon: conservation implications

Populations of many Afro-Palearctic birds have declined, with those wintering in sub-Saharan Africa, such as Wood Warbler Phylloscopus sibilatrix, particularly affected. In this study we investigated the relationship between habitat characteristics and Wood Warbler presence/absence in the Centre Region of Cameroon. A total of six transects were established in three habitat types (forest, forest–savanna transitional zone and savanna). Call playback surveys were conducted monthly from November 2015 to April 2016 to determine Wood Warbler presence/absence. Detailed habitat measurements were also recorded in each transect. A total of 86 responses were recorded: 33 (mean 6.6 ± 2.3) in forest habitat, 47 (mean 9.4 ± 3.36) in the forest–savanna transitional zone, and 6 (mean 2 ± 1.1) in savanna habitat. Wood Warbler presence increased significantly with the number of trees between 3 and 7 m in height, and decreased significantly with the number of shrubs between 0.5 and 3 m in height. Anthropogenic disturbance such as the agricultural cycle and burning were not found to have an effect on Wood Warblers presence/absence. We conclude that Wood Warblers overwinter in all three habitat types with probability of detection greatest in the forest–savanna transitional habitat with a relatively low canopy and an open understorey. Forest clearance in sub-Saharan Africa potentially threatens wintering habitat for Wood Warblers.     

Habitat selection by wapiti (Cervus elaphus xanthopygus) in Wandashan Mountains, China, based on habitat availability

In 2004 and 2005, we studied winter habitat use and selection by wapiti in Wandashan Mountains, Heilongjiang Province, China. Using Geographic Information System (GIS) and Remote Sensing (RS), we measured habitat availability in terms of the area of each habitat category in the study area. We compared winter habitats used by wapiti and their availability. The results showed that the habitats used by wapiti were disproportional to their availability. Wapiti preferred clear-cuttings, low vegetation coverage (No. 2, 3 NDVI classes), 200–300 m elevation range, south aspect, and 600–700 m distance range away from roads. They also showed different preferences for feeding and resting. The feeding sites tended to be located at broad-leaved stands and clear-cuttings, No. 2, 3 NDVI classes, and south slope, whereas the bedding sites tended to be clear-cuttings, and No. 2, 3, 4 NDVI classes. Using the logistic regression model to predict the probability of habitat use by wapiti in winter, we found that the overall prediction accuracy was 74.4% for the total habitat samples, with 84.2% for the habitat samples used by wapiti and 62.4% for the control samples correctly predicted by using this model.