姜寨遗址先民食谱分析

尽管通过C、N稳定同位素分析已对陕西临潼姜寨遗址先民的食物结构和粟作农业进行了初步探索,但研究中的一些重要问题,如不同时期先民食物结构的演变,姜寨与半坡、史家先民食物结构间的差异等,依然未能深入讨论。为此,本文对姜寨遗址出土的不同时期(一期、二期)人骨进行了C、N稳定同位素分析,试图揭示先民食物结构的演变历程,探索姜寨与半坡、史家先民食物结构间的差异,并初步探讨产生这种差异的可能原因。人骨的δ13C平均值(-9.7±1.0‰)和δ15 N平均值(8.5±0.5‰),表明粟类食物(包括粟类作物以及依赖于粟类作物的动物等)在姜寨先民食谱中占据主要地位,且动物资源在食物中的比例相对较低。两期先民的δ13C和δ15N值,无显著性差异,表明先民一直从事粟作农业以及家畜的饲养活动。食谱分析并未发现第二期先民食用更多野生动物的证据,这可能与先民样品量相对偏少、二期先民食用的野生动物比例上升较小、动物类食物在先民食谱中所占比例较小等原因有关。对比地理位置毗邻、文化年代相近的姜寨、史家、半坡遗址先民δ13C值,姜寨(-9.7±1.0‰,N=19)与史家(-10.0±0.7‰,N=9)接近,而远高于半坡(-14.8±1.9‰,N=5),表明半坡遗址粟作农业的种植规模要逊于姜寨和史家遗址。我们认为,不同遗址间先民δ13C值的差异,可能受半坡样品量偏少、遗址间小生态环境不同等因素的影响。     

哈民忙哈遗址人和动物骨骼的C、N稳定同位素分析

哈民忙哈遗址是迄今为止内蒙古乃至东北地区发现面积最大的一处史前聚落遗址,为重建当时的生产和生活情况、文化习俗甚至思想观念等都提供了弥足珍贵的实物资料。其中,重建该遗址先民的食物结构和生业经济,有助于揭示哈民忙哈遗址产生和兴盛的动因及科尔沁沙地史前文化的变迁过程。目前,多学科的研究成果已经初步显示哈民忙哈遗址先民的生业经济具有多样性,但各种生业模式的比重以及是否存在家畜饲养等问题还缺乏相应的了解。本文对该遗址87例人骨与18例动物骨骼进行了C、N稳定同位素分析,还原了人和动物的食物结构及生业经济面貌。结果显示,陆生野生动物的食物结构主要基于C₃植物类食物和少量的C₄农作物,而犬科动物则体现出家养动物的食谱特性：食物主要依赖C₄食物和人类食物残羹。C₄类粟黍农作物和以此饲喂的动物是该遗址先民的主要食物来源,其中粟黍农作物在食物结构中的地位尤其重要。在该遗址中,女性摄入相对较多的粟黍和相对较少的肉类,与男性在食物结构上存在显著差异。农耕、家畜饲养是先民最重要的经济行为,狩猎、渔猎和采集是生业的重要补充。     

新疆巴里坤东黑沟遗址出土人骨的碳氮同位素分析

本文以新疆巴里坤东黑沟遗址出土的人骨为研究对象, 测定骨中的碳、氮同位素比值, 揭示该地区古代先民的食谱特征, 探索当时人们的生活与生产方式、经济形态等历史信息。研究结果发现: 人骨样品的δ13C值分布在-18.96‰~-17.74‰范围内, 样品的δ15N值均较高, 分布在12.64‰~14.33‰范围内, 表明新疆巴里坤东黑沟遗址先民的食物结构是肉类食物在日常饮食生活占有相当大的比例。根据考古发掘出土的动植物遗存, 肉类的获取应来源于驯养的牛、羊等或部分狩猎所得的鹿等动物, 植物性食物主要来源于大麦、小麦等C3类作物, 这与其生活的环境和以游牧为主, 农业和狩猎为辅的经济形态密切相关; 男性与女性的食物结构相同。     

山西乡宁内阳垣遗址先民食物结构分析

本文应用C、N稳定同位素分析方法,对山西乡宁内阳垣遗址先民的食物结构进行了分析,旨在揭示该遗址先民的生活方式及社会经济状况,探索戎狄对该遗址先民的影响。分析表明,该遗址先民的δ13C平均值为-8.27‰,δ15N平均值为9.54‰,由此推测他们主要的经济模式应为畜牧业。考古学证据显示该遗址主体文化为晋,但食物结构分析却反映,先民的经济模式主要为畜牧业,明显受到了游牧民族(戎狄)的影响。样品XNM50具有异常的δ13C值(-15.36‰)和较高的δ15N值(9.88‰),当与其从事狩猎活动有关。δ13C和δ15N的统计分析表明,不同性别的先民在食物结构上无明显差异,而不同规格的墓葬却差异明显。     

辽宁北票喇嘛洞遗址出土人骨稳定同位素分析

喇嘛洞遗址是我国北方地区一处以三燕文化墓葬为主的大型墓地,对其出土人骨进行C和N稳定同位素分析,可望了解先民的食谱,揭示其生活方式进而探索鲜卑与汉的相互联系。该遗址20个墓葬出土人骨的C、N含量以及C/N摩尔比反映了样品均保存较好,可用作食谱分析。人骨中骨胶原δ13C和δ15N分析显示,先民主要以C4类食物为食,且基本为素食,这当与发达的糜子和粟米农业密切相关,表明喇嘛洞遗址鲜卑生活方式已受到汉文化的强烈影响,由游猎为生转变为农业生产为主。对不同性别的先民食谱差异性进行t检验,发现男女食谱上不存在显著差异。样品BLM63和BLM99的δ15N值相对较高,揭示了其食物结构中含有一定量的肉类。     

通过稳定同位素分析云南大阴洞遗址先民的生业经济方式

农业的起源和传播是人类历史上最重要的变革之一。本文通过对云南省文山州广南县大阴洞遗址出土的17例人骨标本的C、N稳定同位素分析,探索新石器时代晚期至青铜时代,滇东南地区先民的食物结构和生业经济。结果显示,大阴洞遗址居民的δ¹³C值范围为-20.2‰ ~ -17.5‰,主要以C₃类食物为主;δ¹⁵N值范围为9.2‰ ~ 10.7‰,所处的营养级别较高;两性之间存在食谱差异,大部分男性比女性摄入更多的动物蛋白。结合相关考古发现,大阴洞遗址居民可能主要经营稻作农业,兼营狩猎;对动物资源的利用可能具有多样性,并通过采集、渔猎及家畜饲养补充食物来源。考虑到云南作为农业向中南半岛传播的重要通道,进一步对云南地区新石器-青铜时代先民的生业策略选择的梳理发现,云南地区新石器时代不同区域的人群有不同的农业经营策略。到青铜时代,随着小麦的传入,农业经济的选择更为多元化,动物资源的获取技术也更加成熟多样。不同的农业策略选择除了受到自然条件的影响,也是与相邻地区文化互动的结果。     

甘肃西山遗址早期秦人的饮食与口腔健康

本文以甘肃礼县西山遗址早期秦人的遗骸资料作为主要研究对象,以牙齿磨耗、龋病、生前牙齿脱落、上下颌骨表面骨质隆起等指标作为主要观察内容,与国内外相关样本组作对比分析;与此同时,结合食谱分析结果,西山先民饮食状况较为复杂多样,总体上以肉食居多,植物性食物也占一定比例;推测早期秦人的经济模式应为农牧兼营的混合经济模式.     

相近社会等级先民的食物结构差异——以山西聂店遗址为例

一般认为,先民食物结构的差异是人类不同社会等级的重要表现形式之一。然而,对于考古学资料中未见明显等级差异的先民来说,其是否存在食物结构上的差别,仍需认真加以探讨。为此,本文对山西聂店遗址墓葬(基本无随葬品或随葬品较少)中出土的人骨进行了C、N稳定同位素分析,研究结果表明:先民的食物结构,体现了典型中国北方农业经济的特点,即以粟作农业和家畜饲养为生。此外,尽管在考古学意义上聂店先民的等级相近,但其在食物资源的获取上却显示出明显差异,且这种差异与先民的性别、年龄和随葬品的种类和多寡无关。因此,在社会等级较为模糊的人类社会,先民对食物资源的获取,很可能更多地受到人群组成、生活习惯等因素的影响,而与社会等级的相对高低无关。     

广东湛江鲤鱼墩遗址人骨的C、N稳定同位素分析:华南新石器时代先民生活方式初探

相对于长江流域和黄河流域而言,华南地区先民生活方式的了解一直较为缺乏。广东湛江鲤鱼墩遗址人骨的C和N稳定同位素分析,显示此遗址的先民主要以海生类资源为食,陆生资源(包括可能的块茎类原始农业和动物)只占次要地位。通过与约同时代黄河流域、长江流域先民的稳定同位素数据比较,可以看出,早在6000年前,我国先民的生活方式就已出现3大类型。     

通过烟台午台遗址人牙结石淀粉粒探究人类对植物资源的利用

本研究通过对山东烟台市午台遗址人牙结石淀粉粒的分析,发现了来自禾本科的粟（Setaria italica L.）、小麦族（Triticeae）、壳斗科栎属（Quercus sp.）、豆科（Fabaceae）及块根块茎类植物的淀粉粒,数量较为丰富,种属来源较为多样,显示了午台遗址先民利用植物性食物资源的多样性。研究表明午台先民除了利用粟黍类农作物作为主要食物资源外,还广泛采集一些野生果类和块根块茎类作为食物的补充。本研究中发现了大遗存中没有发现的壳斗科栎属等,弥补了大遗存研究的不足。     

河南舞阳张王庄遗址仰韶早期先民食物的淀粉粒分析

黄淮西部是我国新石器时代南、北不同考古学文化与农业模式的过渡地带,近年来该地区植物考古研究成果显著,但目前学术界对该区域仰韶早期人类植食资源利用与农业发展状况仍不甚了解。本文利用淀粉粒分析方法,对张王庄遗址出土仰韶文化早期54件陶器和13件石器进行了表面残留物提取与分析。结果显示,该遗址仰韶早期先民的植食资源利用具有显著的多样性特征,旱地农作物粟、黍虽然已在人类生业经济中占据了重要地位,但采集获取的各类野生植物资源包括薏苡属、小麦族、莲藕等仍是人类食物主要组成,其重要性甚至高于农业种植。与此同时,研究结果也表明,在至迟不晚于距今6000年的仰韶早期阶段,粟、黍两类旱地作物已传播至黄淮西部的低纬度地区并成为先民种植的主要农作物,从而在整个黄淮西部形成以粟、黍为主,水稻为辅的稻-粟兼作农业。研究结果首次提供了黄淮西部仰韶早期人类植食资源利用与农业发展状况的科学证据,对认识黄淮西部史前农作物传播与农业结构演化的具体时空过程等具有重要价值。     

Quantitative analysis of the use of discards from squid trawlers by Black-browed Albatrosses Diomedea melanophris in the vicinity of the Falkland Islands

In the past decade, a major trawl fishery for the squid Loligo gahi has developed in the vicinity of Beauchêne Island, an internationally important breeding site for the Black-browed Albatross Diomedea melanophris. The breeding season diet of this albatross in the Falklands and its use of discards generated by the Loligo fishery were investigated. Albatross chicks are fed extensively on commercially exploited species of squid and fish including Loligo gahi and southern blue whiting Micromesistius australis. The quantity of waste generated by the Loligo fishery amounts to c. 5% of the reported catch and just over 50% of this waste, mainly Loligo and nototheniid fish, is scavenged by adult Black-browed Albatrosses. The total quantity scavenged during the chick rearing period amounts to 1000–2000 tonnes per year. This is equivalent to 10–15% of the total food requirement of the breeding Black-browed Albatross population on Beauchene Island during the period when the fishery is operating. Although the Loligo fishery currently provides a significant quantity of food to these albatrosses, its net effect may be detrimental to them, as it is a much greater predator of Loligo stocks than the albatrosses are estimated to have been prior to the fishery's development.     

The European badger Meles meles: food specialist or generalist?

Although most authors characterize the badger Meles meles as an omnivore, the species is sometimes described as a food specialist and in particular as a specialist predator on earthworms Lumbricus terrestris . The validity of the'food specialist'hypothesis is examined, using data from 11 quantitative studies of badger diet to test two predictions: 1) any one population of badgers relies to a disproportionate extent on a single type of food; and 2) consumption of the predominate food is independent of its availability. As regards the first prediction, when data on year-round consumption were corrected for seasonal variation in total intake, no study showed earthworms to comprise more than 50% of the diet and in three studies they accounted for no more than 5% of intake. In two studies, fruit comprised more than 50% of the diet and fruit and insects together accounted for 95–97% of total consumption. Compared across different studies, consumption of any one food, including earthworms, varied more or less continuously across a wide spectrum of values, providing no support for the idea that a given food can be characterized as being of either major or minor importance. With respect to the second prediction, there was no evidence that consumption of either earthworms or any other food was consistently invariant across seasons; and in any case, the idea that badger diet contains a constant proportion of any one food does not make functional sense. Taken together with information about the morphology of the digestive system and a priori arguments about the conditions in which food specialization is likely to be adaptive, studies of diet confirm that the badger is best viewed as a generalist or opportunist feeder.     

Landscape-mediated variation in diet is associated with egg size and maculation in a generalist forager

Human impacts alter landscapes with consequences for the distribution and availability of high-quality food resources to populations inhabiting those landscapes, which may impact on the reproductive output of individuals in those populations. The sensitivity of wild populations to changes in food resources may vary among stages of the annual cycle. For example, in birds, effects are likely to be greater during costly stages such as egg production. Here we compare assimilated diet (from stable isotope analysis of chick feathers) and egg traits (egg size, shape, eggshell colour and maculation, using pattern-analysis software) in Herring Gulls Larus argentatus, across seven colonies in southwest Scotland and Northern Ireland. The Herring Gull is an opportunistic, generalist forager on both marine and terrestrial resources which frequently exploits anthropogenic food sources such as fishery discards and human refuse. We found that larger eggs were laid in colonies where females consumed either a higher proportion of marine resources or terrestrial resources; smaller eggs were laid in colonies where females had an intermediate diet. In colonies where females consumed more marine items, they also laid eggs with higher maculation (intensity and size of spots) compared with colonies where females mainly consumed terrestrial food. We also found smaller and more pointed eggs, suggestive of resource shortages, in larger colonies. Generalist foragers are often thought to have the capacity to buffer themselves against changes in the food web, provided that enough alternative food is available. However, this study highlights that specializing on the most profitable or available resources has consequences for egg traits even in an opportunistic generalist forager exploiting a large range of habitats. If variation in egg traits is related to reproductive output, then understanding the impact of assimilated diet on reproduction early in the breeding season can provide important insights into how populations will respond to landscapes altered by human impact.     

珠江口底栖动物生态学研究

采泥样调查所获底栖动物一般为体形较小,活动能力较弱的种类,往往是一些底层鱼类和虾类等良好的天然饵料。珠江口海域渔业资源丰富,是中国南海区的主要渔场之一,因此,对该海域饵料底栖动物进行研究有重要意义。通过1999年9月和2000年4月分别对珠江口底栖生物进行的两个航次的采泥样调查,分别鉴定出底栖动物15衙睡21种,共计32种。底栖动物的优势种,秋季为光滑河篮蛤Potamocorbula laevis(Hinds)(Y=0.387),春季为光滑河篮蛤（Y＝0．464）和欧虫Owenia fusformis Delle Chiaje(Y=0.120)。平均个体数量和生物量,春季为591.7ind./m^2和26.7g/m^2,秋季为85.0ing./m^2和7.4g/m^2;而在各类群生物中,软体动物占绝大部分,其次为多毛类,其它各类群所占比例都不足5％。生物多样性,个体数量和生物量的分布均呈由北向南增加的趋势。另外,与近20a中的历史资料相比,珠江口底栖动物个体数量变化不大,除2000年春季较高为591.7ind./m^2外,其变化范围在72.4-128ing./m^2之间。春季生物量除1991年较高为27．0g/m^2外,变化不是很大,大约10g/m^2;但秋季生物量呈明显下降趋势,1980年为30.1g/m^2,1990年为27.8g/m^2,1999年秋季急剧下降至7.4g/m^2。此外,从底栖动物各大类群的百分组成变化情况来看,一般以软体动物个体数量（22．7％－83．2％）和生物量（57．9％－82．5％）都最高;多毛类的个体数量百分组成占第二倍,其范围是13．4％－52．3％;其它各类群的百分组成除个别时候所占比例较大外,一般都较小。     

Contribution of allochthonous resources to breeding in a high-arctic avian predator

Allochthonous input of resources (i.e., originating from a place other than where they are found) can have a significant impact on food availability for consumers. We assessed the impact of an allochthonous source of food (the sewage outfall stream of a military base) on an avian predator breeding in a low-productivity, high-arctic site (Alert, 83°N, 62°W, Ellesmere Island), the long-tailed skua (Stercorarius longicaudus). We collected blood samples throughout the breeding season during two contrasting years of lemming abundance to characterize diet composition of skuas and evaluated the contribution of the anthropogenic and lemming food sources using stable isotopes (carbon ¹³C and nitrogen ¹⁵N). The isotopic signature of skuas changed seasonally because they switch from a marine to a terrestrial diet when they come ashore to breed but also differed between the 2 years of the study. Anthropogenic food source accounted for 33 % of the summer diet but this proportion varied between years, from 41 % (5–95 ‰: 13–59 %) in a year of low lemming abundance to 16 % (5–95 ‰: 10–21 %) in a high year. Skua nest density recorded in years of low lemming abundance at Alert (0.15 nests/km²) was higher than at any other comparable arctic sites (0–0.02 nests/km²). Overall, the use of an anthropogenic food sources apparently subsidizes skua reproduction at this site, which could affect the food web of this low-productivity ecosystem.     

黄陵寨头河与史家河墓地人骨的碳氮稳定同位素

戎人在古代多样性文化历史发展进程中有着不可忽视的作用,由于文献记载的匮乏,对其生活历史信息的认知极少。陕西延安黄陵寨头河与史家河战国时期戎人墓地的考古发掘,为了解古代关中地区戎人的社会生活提供了珍贵的实物资料。本文则以两墓地出土的人骨为研究对象,采用C、N稳定同位素分析方法,讨论先民的饮食结构,探索其经济模式。结果表明：寨头河与史家河墓地先民的食谱相似,食谱中植物性食物主要来自粟、黍等C₄类作物;除史家河墓地两个个体外,其余先民动物蛋白摄入较少,且未因性别不同而产生食谱差异。史家河墓地战国晚期秦人与早中期戎人的饮食结构相似。结合其他考古资料以及文献可知,寨头河与史家河墓地先民吸收关中农业文化后,利用当地良好的自然环境发展农业,形成了以农业种植为主、农牧混合的经济模式。     

The human adaptations to meat eating: a reappraisal

In this paper we discuss the hypothesis, proposed by some authors, that man is a habitual meat-eater. Gut measurements of primate species do not support the contention that human digestive tract is specialized for meat-eating, especially when taking into account allometric factors and their variations between folivores, frugivores and meat-eaters. The dietary status of the human species is that of an unspecialised frugivore, having a flexible diet that includes seeds and meat (omnivorous diet). Throughout the various time periods, our human ancestors could have mostly consumed either vegetable, or large amounts of animal matter (with fat and/or carbohydrates as a supplement), depending on the availability and nutrient content of food resources. Some formerly adaptive traits (e.g. the “thrifty genotype”) could have resulted from selective pressure during transitory variations of feeding behaviour linked to environmental constraints existing in the past.