北极-第三纪成分在喜马拉雅-横断山的发展及演化

通过古植物学及古地史资料,论述了北极地区在第三纪早期气候变迁对喜马拉雅-横断山地区植物区系形成的影响,并通过一些喜马拉雅-横断山地区的特征类群;杜鹃属,槭树属,柳属,桦木属和桤木属以及一些草本类群虎耳草科的落新妇族,金腰属和虎耳草属,忍冬科的莛子镳属的分布式样和起源分化的分析,说明了喜马拉雅-横断山区系中有相当一部分是这些北方起源的,北极-第三纪成分迁移是现代喜马拉雅-横断山植物区系成分的一个重要来源,并且讨论了喜马拉雅-横断山植物区系同北方植物区系可能的迁移路线,指出西南-秦岭-东北通道即西南沿四川盆地经秦岭和黄河至东北和西伯利亚和“中亚高地通道”即向西经帕米尔高原同北方交流,向东经西南-秦岭-东北通道迁移交流是喜马拉雅-横断山植物区系成分同北方交流的主要路线,由于第四纪冰期间冰期反复作用,各类成分在该地区保存,汇集,分化发展,使得该喜马拉雅-横断山地区成了北极-第三纪中心消失以后,在近代形成的温带和高山植物区系新的发展和分化中心。     

青藏高原鱼类区系特征及其形成的地史原因分析

本文以作者大量实地考察资料为基础,对青藏高原鱼类区系特征进行深入分析和归纳,提出青藏高原鱼类6科26属114种的名录及各水系间鱼类共有属种的比较图。首先阐述了青藏高原鱼类区系组成的单纯性与复杂水系的概况,进而找出青藏高原与新疆等干旱地区间的鱼类区系和高原内部各水系间的鱼类区系的相互联系。第二部分,通过分布特征的分析确定青藏高原鱼类分布的三大特点,即在外泄河流上有明显的地理分界;在东南外围水域中呈点状分布,在垂直高度上有明显的地带性。最后,基于区系特征的分析,联系有关古地理、古气候和鱼类化石资料,探讨了形成青藏高原鱼类区系的地史原因。认为适应于温暖水域生活的原始鲃类和鳅类,在早第三纪时已广布于东亚和中亚地区。青藏高原现代鱼类区系不是由一个起源中心辐射或跨阻传布所形成的,而是通过喜马拉雅运动的三次重大地质历史事件产生隔离,导致青藏高原鱼类与其它地区鱼类的分离而形成的,其分别是:1.喜马拉雅地槽闭合引起“西康群山”再次隆起构成中亚与东亚鱼类区系的分离;2.喜马拉雅山隆起引起中亚与南亚鱼类区系间分离;3.阿尔金山等山脉再度隆起构成中亚地区青藏高原鱼类区系的分离,最后完成青藏高原这一独特的鱼类区系的雏形。 此外也论述了青藏高原鱼类区系东部分界线实际     

甘肃省马衔山地区种子植物区系的研究

分析了甘肃省马衔山地区种子植物区系,用主成分分析（Ｐ．Ｃ．Ａ）和信息聚类（Ｉ．Ｃ．Ａ）方法对该区系与国内其它１０个有代表性地区的种子植物区系进行了对比分析。其结果为：１．该区系温带性质十分明显,是一个汇集四大区系成分的过渡带;２．该区系起源于第三纪亚热带亚高山森林植物区系,随青藏高原的隆起和中亚干旱区的形成而分化形成;３．该区系与六盘山植物区的关系最近。该区系隶属于泛北极植物区,中国－日本森林植物     

中国菊科植物的系统分类与区系的初步研究

为1993年"菊科植物的系统分类与区系地理的初步探讨"(世界)一文的姊妹篇,重点论述我国菊科的系统分类及其区系地理成分。文中介绍了分布我国的菊科240属隶于2亚科、5超族、11族中的系统位置。论述了我国菊科植物区系地理成分的特点是:1.大洲间共同分布或洲际间断分布的属多,且具明显的热带亲缘;2.与亚洲国家,包括中亚国家或亚洲热带国家共同分布的属多,尤其是成"中亚-青藏高原-喜马拉雅山"地区分布的属多;3.中国特有属多,其中我国西南省区特有属最多。文中还讨论了分布我国菊科各族祖先种的起源、迁移以及我国区系地理热带亲缘和"横断山脉-喜马拉雅山脉(东)森林植物亚区"菊科植物分布的特点。     

鹅观草属的地理分布

鹅观草属是禾本科小麦族中的最大的属,现知全世界有4组,20系,126种,分布于北半球的温带和寒带,中国有4组,18系,79种,主要分布于西北,西南,华北和东北,是鹅观草属植物种类最为集中的区域,尤其高原东北部的唐古特地区又是我国鹅观草属分布相对密集之地,有3组,12系,30种,而且其间不同等级,不同演化水平的类群均有分布,该地可能就是该属的现代分布中心,同时,唐古特地区多汇聚有鹅观草属不同等级的原始类群和与原始类群有很缘的短柄草属植物,其中最原始的大柄鹅观草特产于该区,而该区缺乏的是高级的大颖组类群,故推测唐古特地区可能又是该属的起源地,起源时间大约在青藏高原明显增高,气候转凉的晚第三纪初的中新世,鹅观草属起源后,在中国境内地质活动比较剧烈的地区得到了进一步的发展和分化,但只有少数适应性较强的类群大概以3条路径扩展到国外,并向东到在北美的巴芬岛,向西延伸到大西洋滨岸,向北进入寒冻的北极地区。     

藜科植物的起源、分化和地理分布

全球藜科植物共约130属1500余种,广泛分布于欧亚大陆、南北美洲、非洲和大洋洲的半干旱及盐碱地区。它基本上是一个温带科,对亚热带和寒温带也有一定的适应性。本文分析了该科包含的1l族的系统位置和分布式样,以及各个属的分布区,提出中亚区是现存藜科植物的分布中心,原始的藜科植物在古地中海的东岸即华夏陆台(或中国的西南部)发生,然后向干旱的古地中海沿岸迁移、分化,产生了环胚亚科主要族的原始类群;起源的时间可能在白垩纪初,冈瓦纳古陆和劳亚古陆进一步解体的时期。文章对其迁移途径及现代分布式样形成的原因进行了讨论。     

大理苍山种子植物区系的研究

通过对苍山种子植物164科,852属,2 503种,45亚种,194变种在各个层次不同角度上的分析,探讨了本区种子植物区系的性质、地位、起源及演化。区系分析表明:大理苍山种子植物区系在我国区系区划上属于东亚植物区,中国-喜马拉雅森林植物亚区,横断山脉地区,三江峡谷亚地区;其区系具有鲜明的温带性质;现代苍山植物区系是在印度板块与欧亚板块碰撞,古地中海退却和喜马拉雅隆升等地质事件背景下,由古地中海成分、北极—第三纪成分及古南大陆成分融汇发展而来;苍山的特有现象较为显著,新老皆备,且以新特有成分为主,它既是古特有成分的避难所,也是新特有成分发展的温床;苍山在植物分布上是呼应东、西、南、北的纽带,同时又是很多成分分布的边界,是重要的区系节。     

青藏高原和喜马拉雅地区锦鸡儿属植物的地理分布

锦鸡儿属Caragana是一个典型的温带亚洲分布属。本属在青藏高原和喜马拉雅约有24种1变种,约占整个属的1／3。这些种类几乎全部处于演化高级阶段,且既有叶轴宿存类群,也有假掌状叶类群。反映出种的分化很活跃,在横断山地区形成本属的分布中心、分化中心。本区内绝大多数种类是特有分布。替代现象主要受气候、植被变化作用,沿横断山和喜马拉雅分布的长齿系Ser. Bracteolatae Kom．是一个典型的替代分布类群。锦鸡儿属植物生态适应性很强,可在其生长的灌丛中形成优势种。 寒化和旱化现象十分突出,它们有一系列森林种、草原种和荒漠种及相关的形态变异。用锦鸡儿属植物进行青藏高原和喜马拉雅区域内的分布区关系分析及最小生成树MST和特有性简约性分析(PAE),表明横断山地区特别是其北部是本属植物的一个地理结点。以此沿横断山向北部唐古特和西部藏东南适应性辐射。横断山和西喜马拉雅联系微弱,看不出植物长距离扩散的踪迹,大多是由于生态因子限制而产生的隔离。虽然本区不可能是锦鸡儿属的起源地,然而,通过本区与邻近地区的地理联系,可推测它们在我国适应性辐射方向是从东北向西南。结合豆科蝶形花亚科其它属化石记录及其分布区局限在温带亚洲等现象,认为锦鸡儿植物是一组特化、晚近衍生的类群,起源于北方东西伯利亚晚第三纪中新世后期至上新世。     

青藏高原特有种子植物区系特征及多样性分布格局

青藏高原拥有丰富的种子植物, 但对该地区特有植物的区系特征以及多样性还鲜有报道。本文通过植物志(书)以及在线数据库, 整理了只分布于青藏高原地区的种子植物名录及其地理分布, 分析了它们的科属特征、区系成分以及多样性空间分布格局。结果表明: 青藏高原共有特有种子植物3,764种, 隶属113科519属, 多数为草本植物(76.3%); 包含100种以上的科有菊科、毛茛科、列当科等15个, 属有马先蒿属(Pedicularis)、杜鹃花属(Rhododendron)、紫堇属(Corydalis)等7个; 从属的区系成分来看, 温带成分占主导(67.5%)。青藏高原特有植物多样性格局呈现从高原东南部向西北部逐渐递减的趋势, 其中东喜马拉雅-横断山脉的物种多样性非常丰富, 而且多数物种分布在高原的中海拔地带。理解青藏高原特有物种的特征及多样性格局对探讨高原植物区系的演化历史和物种保护有重要启示。     

豆科黄华属的植物地理研究

首次全面论述了全世界黄华属(豆科)植物地理。黄华属是豆科少数几个东亚-北美间断分布属之一。对黄华属5组21种的分布进行了分析,发现本属4个频度分布中心依次是：东亚地区(8种／3组,其中特有种4种),伊朗-土兰地区(7种／3组,其中特有种3种),落基山地区(7种／2组,均为特有种)及大西洋北美地区(3种／1组,均为特有种)。基于以下事实：在东亚地区存在本属最多的组与种;在此区可以见到黄华属系统发育系列;该属最原始的组种及最进化的组种也在该区出现等,可以认为东亚地区是该属的现代分布中心及分化中心。伊朗-土兰地区(中亚东部至喜马拉雅)及落基山地区所含种、组数仅次于东亚地区,而且多倍体现象多发生于这两区,因此可认为是本属的次生分布中心及分化中心。在此二地区,物种分化较活跃且复杂,先后描述了很多新种和变种,也曾进行过较多的归并处理。最近的分子生物学证据不断揭示,在这地区曾被归并的一些分类群存在着较大不同,从而提醒分类学家对年轻区系中物种分化较活跃的类群进行分类处理时,无论是建新分类群还是对某些类群进行归并,应持谨慎态度。作者根据黄华属植物的现代地理分布、形态演化趋势、现有的化石及地质历史资料,推测黄华属植物在中新世之前早已形成,并且在晚第三纪欧亚大陆与北美大陆失去陆地连接之前在两大陆已经存在,很可能是于早第三纪或晚白垩纪在劳亚古陆上起源于一个含羽扇豆生物碱的古槐成员。两大陆分离后,在不同的成种因子的影响下,形成了各自的演化格局：在亚洲,晚第三纪的喜马拉雅造山运动、古地中海消失及第四纪冰川作用引起的旱化、寒化,促进了该属植物的强烈分化;而在北美,第四纪的冰川作用及局部的山体隆起,可能是促进该属植物演化的主要动力。根据黄华属植物的系统演化趋势及原始类群的分布式样分析,东亚地区的中国-日本亚区可能是本属植物的原始类型中心。     

中国红原泥炭沼泽植物区系研究

沼泽植物是形成泥炭物质来源的重要资源植物。我国省西北部高原的红原县是我国沼泽最发达的地区之一,沼泽植物约有１５５种,隶属于３９科,８７属,包括世界分布属２０属,泛热带分布属５１属,旧世界温带分带属７属,温带亚洲分布属２属,地中海,西亚至中亚分布属１属和东亚分布属５属,在泥炭沼泽植物中,莎草科是种数最多的科,其次是菊科、毛茛科、玄参数、禾本科和龙胆科,它们的种数约占全部泥炭沼泽植物种数的５３．５４％     

The Compositae of Xizang (Tibet)

1) The Compositae in Tibet so far known comprise 508 species and 88 genera, which nearly amounts to one fourth of the total number of genera and one third of the total number of species of Compositae in all China, if the number of 2290 species and 220 genera have respectively been counted in all China. In Tibet there are all tribes of Compositae known in China, and surprisingly, the large tribes in Tibetan Compositae are also large ones in all China and the small tribes in Tibet are also small ones in all China. Generally speaking, the large genera in Tibet are also large ones in all China and the small genera in Tibet are likewise small ones in all China. In this sense it is reasonable to say that the Compositae flora of Tibet is an epitome of the Compositae flora of all China. In the Compositae flora of Tibet, there are only 5 large genera each containing 30 species or more. They are Aster, Artemisia, Senecio, Saussurea and Cremanthodium. And 5 genera each containing 10—29 species. They are Erigeron, Anaphalis, Leontopodium, Ajania, Ligularia and Taraxacum. In addition, there are 77 small genera, namely 87% of the total of Compositae genera in Tibet, each comprising 1—9 species, such as Aja-niopsis, Cavea and Vernonia, etc. 2) The constituents of Compositae flora in Tibet is very closely related to those of Sichuan-Yunnan provinces with 59 genera and 250 species in common. Such a situation is evidently brought about by the geographycal proximity in which the Hengtuang Shan Range links southeastern and eastern Tibet with northern and northwestern SichuanYnnnan. With India the Tibetan Compositae have 59 genera and 132 species in common, also showing close floristic relationships between the two regions. Apparently the floristic exchange of Compositae between Tibet and India is realized by way of the mountain range of the Himalayas. The mountain range of the Himalayas, including the parallel ranges, plays a important role as a bridge hereby some members of the Compositae of western or northern Central Asia and of the northern Africa or of western Asia have migrated eastwards or southeastwards as far as the southern part of Fibet and northern part of India, or hereby some Compositae plants of eastern and southeastern Asia or Asia Media have migrated northwestwards as the northern part of Central Asia. Some of the species and genera in common to both Tibet and Sinjiang indicate that this weak floristical relationship between these regions is principally realized through two migration routes: one migration route is by way of the Himalayas including the parallel ranges to Pamir Plataeu and Tien Shan, or vice versa. The other migration route is by way of northern Sinjiang to Mongolia, eastern Inner Mongolia, southwards to Gansu, Qinghai (or western Sichuan), eastern Tibet up to the Himalayas, or vice versa. However, Tibet is not entirely situated at a migration crossroad of the floral elements. An ample amount of the data shows that Compositae flora have a particular capability of development in Tibet. of the total number of species of Tibetan Compositae, 102 species and 1 genus (Ajaniopsis Shih) are endemic. Besides, 8 genera are regional endemics with their range extending to its neighbourhood. The higher percentage of endemics at specific level than at generic in Tibetan Compositae may be a result of active speciation in response to the new enviromental conditions created by the uplifting of the Himalayas. The flora in Tibetan Plateau as a whole appears to be of a younger age. 3) The uprising of the Himalayas and of the Tibetan Plateau accompanied by the ultraviolet ray radiation, the microthermal climate and the high wind pressure has, no doubt, played a profound influence upon the speciation of the native elements of Tibetan Compositae. The recent speciation is the main trend in the development of the Com-positae flora native in Tibet in the wake of upheaval of the plateau.     

On the geographical distribution of the Juglandaceae

The present paper aims to discuss the geog raphical distribution of the Juglandaceae on the basis of unity of the phylogeny and the process of dispersal in the plants. The paper is divided into the following three parts: 1. The systematic positions and the distribution patterns of nine living genera in the family Juglandaceae (namely, Engelhardia, Oreomunnea, Alfaroa, Pterocarya, Cyclocarya, Juglans, Carya, Annamocarya and Platycarya) are briefly discussed. The evolutional relationships between the different genera of the Juglandaceae are elucidated. The fossil distribution and the geological date of the plant groups are reviewed. Through the analysis for the geographical distribution of the Juglandaceous genera, the distribution patterns may be divided as follows: A. The tropical distribution pattern a. The genera of tropical Asia distribution: Engelhardia, Annamocarya. b. The genera of tropical Central America distribution: Oreomunnea, Alfaroa. B. The temperate distribution pattern c. The genus of disjunct distribution between Western Asia and Eastern Asia: Pterocarya. d. The genus of disjunct distribution between Eurasia and America: Juglans. e. The genus of disjunct distribution between Eastern Asia and North America: Carya. f. The genera whose distribution is confined to Eastern Asia: Cyclocarya, Platycarya. 2. The distribution of species According to Takhtajan’s view point of phytochoria, the number of species in every region are counted. It has shown clearily that the Eastern Asian Region and the Cotinental South-east Asian Region are most abundant in number of genera and species. Of the 71 living species, 53 are regional endemic elements, namely 74.6% of the total species. The author is of the opinion that most endemic species in Eurasia are of old endemic nature and in America of new endimic nature. There are now 7 genera and 28 species in China, whose south-western and central parts are most abundant in species, with Province Yunnan being richest in genera and species. 3. Discussions of the distribution patterns of the Juglandaceae A. The centre of floristic region B. The centre of floristic regions is determined by the following two principles: a. A large number of species concentrate in a district, namely the centre of the majority; b. Species of a district can reflect the main stages of the systematic evolution of the Juglandaceae, namely the centre of diversity. It has shown clearly that the southern part of Eastern Asian region and the northern part of Continental South-east Asian Region (i.c. Southern China and Northern Indo-China) are the main distribution centre of the Juglandaceae, while the southern part of Sonora Region and Caribbean Region (i.c. South-western U.S.A., Mexico and Central America) are the secondary distribution centre. As far as fossil records goes, it has shown that in Tertiary period the Juglandaceae were widely distributed in northern Eurasia and North America, growing not only in Europe and the Caucasus but also as far as in Greenland and Alaska. It may be considered that the Juglandaceae might be originated from Laurasia. According to the analysis of distribution pattern for living primitive genus, for example, Engelhardia, South-western China and Northern Indo-China may be the birthplace of the most primitive Juglandaceous plants. It also can be seen that the primitive genera and the primitive sections of every genus in the Juglandaceae have mostly distributed in the tropics or subtropics. At the same time, according to the analysis of morphological characters, such as naked buds in the primitive taxa of this family, it is considered that this character has relationship with the living conditions of their ancestors. All the evidence seems to show that the Juglandaceae are of forest origin in the tropical mountains having seasonal drying period. B. The time of the origin The geological times of fossil records are analyzed. It is concluded that the origin of the Juglandaceae dates back at least as early as the Cretaceous period. C. The routes of despersal After the emergence of the Juglandaceous plant on earth, it had first developed and dispersed in Southern China and Indo-China. Under conditions of the stable temperature and humidity in North Hemisphere during the period of its origin and development, the Juglandaceous plants had rapidly developed and distributed in Eurasia and dispersed to North America by two routes: Europe-Greenland-North America route and Asia-Bering Land-bridge-North America route. From Central America it later reached South America. D. The formaation of the modern distribution pattern and reasons for this formation. According to the fossil records, the formation of two disjunct areas was not due to the origin of synchronous development, nor to the parallel evolution in the two continents of Eurasia and America, nor can it be interpreted as due to result of transmissive function. The modern distribution pattern has developed as a result of the tectonic movement and of the climatic change after the Tertiary period. Because of the continental drift, the Eurasian Continent was separated from the North American Continent, it had formed a disjunction between Eurasia and North America. Especially, under the glaciation during the Late Tertiary and Quaternary Periods, the continents in Eurasia and North America were covered by ice sheet with the exception of “plant refuges”, most plants in the area were destroyed, but the southern part of Eastern Asia remained practically intact and most of the plants including the Juglandaceae were preserved from destruction by ice and thence became a main centre of survival in the North Hemisphere, likewise, there is another centre of survival in the same latitude in North America and Central America. E. Finally, the probable evolutionary relationships of the genera of the Juglanda-ceae is presented by the dendrogram in the text.     

青藏高原东缘28种风毛菊属植物花粉形态研究

通过扫描电镜对青藏高原高寒草甸风毛菊属28种多年生草本植物的花粉进行了形态观察和比较,结果显示,该属植物花粉形态较为一致。花粉近球形或长球形,极面观三裂圆形,三孔沟,外壁表面具刺状突起。刺间纹饰可分为4种类型:穴状、颗粒状、网状—颗粒状和网状。花粉形态没有大的差异,但刺间表面纹饰、萌发沟的宽窄、刺的大小、分布密度和花粉大小在种间存在差异,可作为种的分类依据。花粉特征聚类分析结果表明,此28种风毛菊属植物花粉大致划分为明显的5个类群,所包含的种分别隶属于风毛菊属的四亚属:雪兔子亚属、雪莲亚属、附片亚属和风毛菊亚属。     

河北省侧蒴藓类植物的地理成分及其与邻近地区的比较

通过对河北省侧蒴藓类植物较详尽的野外考察、标本采集、鉴定和系统研究,已知河北省侧蒴藓类植物17科,66属,196种(包括种下单位),其地理成分可划分为9种类型,北温带分布型占主导地位,东亚分布型次之,热带分布型不起主要作用。运用物种丰富度、优势科比较、属相似性系数等统计方法,对河北与邻近地区的区系地理成分进行了比较、分析和讨论。从河北省与我国东北、内蒙古、山东、秦岭、西藏和横断山等6个地区侧蒴藓类区系组成的比较分析得出,本省该类群物种丰富度位居第4,横断山居于第1位,河北与其它6个地区的共有优势科前3位为青藓科、柳叶藓科、灰藓科;河北与内蒙古、山东相似程度最高,与东北和秦岭相似程度次之,与西藏、横断山相差最大。同时对这7个地区的具体地理成分也作了比较。这些结果有力地说明了河北省侧蒴藓类植物区系具有明显的温带性质,同时兼有浓厚的东亚区系色彩,也表明了中国华北与日本、欧洲和北美东北部在苔藓植物区系物种起源上有深厚的历史渊源。     

中国松科植物的分布型和区系分析

简介松科研究的新进展,根据新近资料划分中国松科植物10属95种的分布型和进行区系分析,结果是北温带分布4属,东亚-北美间断分布2属,中国特有3属和越南-华南分布1属。25个种分布型分属于欧亚温带、东亚和亚洲热带3个植物区域。比较详细分析其4点主要特征;分布的区域差异和不平衡,高的特有性,间断分布和替代分布等。     

甘肃藜科植物研究

根据对藜科植物标本的收集、整理和系统鉴定,本地区藜科植物共有67种,隶属于21属,8族,在我省有2个分布丰富区：河西走廊地区和青藏高原东缘的甘南地区。分析表明,本区藜科植物可划为5个分布型和3个变型,其中以地中海区、西亚和中亚分布最多（占33.4％）,除世界分布和中国特有外,均为温带性地理成分（占71％）,结合对孑遗属的分析,说明了本区系的主要特征为典型的温带性质和具有一定的古老性,并与相邻地区有一定的联系;本区系起源于白垩纪至第三纪的古地中海沿岸,伴随着青藏高原的隆起、海浸海退以及气候的变迁形成了现今的区系成分。根据植物区系分区的原则和方法,将甘肃藜科植物区系划分为5个区系小区：走廊小区,祁连山小区,中部小区,甘南小区和陇南小区,其中走廊小区和甘南小区不仅是本区藜科植物主要的分布区,而且也是我省重要的农业区和畜牧业基地,因此,对各小区的区系特征进行了论述并提出了相应的生产实践的建议,以期为我省的防风固沙、植被恢复和草场建设等提供理论依据。     

陕北黄土高原蒿属植物的分类与分析

陕北黄土高原有蒿属植物30种1变种,居该地产种子植物属中所含种数的首位。所产蒿属植物在不同的植被带中梯度变化明显,替代现象显著。在生态类型上,旱生类型从南向北递增,中生类型从南向北递减。在区系组成上可分为6种分布区类型,即:我国特有分布,3种;温带亚洲分布,14种1变种;北温带及中亚分布各4种;旧世界温带分布,3种;东亚分布的2种。可见陕北黄土高原蒿属植物种类丰富,梯度变化明显,旱化现象显著,地理成分复杂,但以温带亚洲分布类型为主,兼有其它成分,属典型的温带性质。     

广东裸子植物区系的特点

广东省裸子植物共有８科１８属３４种,分别占中国同类的８０．０％,５２．９％和１７．６％。其中泛热带分布２属,热带亚洲至热带大洋洲１属,热带亚洲１属,北温带５属,东亚和北美间断分布１属,东亚分布３属,中国特有分布５属,即热带成分共仅４属占２２．２％,而亚热带至温带成分１４属占７７．８％为绝对优势。分析表明：广东裸子植物区系体现了东亚裸子植物区系的特点,其原始中心和现代分布中心都在中国亚热带。同时论文还将广东裸子植物种的分布区类型划分为２０个亚型。