湖北松滋、建始地区中、上奥陶统牯牛潭组至宝塔组牙形类及其地层意义

本文针对湖北松滋响水洞剖面及建始小茶园剖面牯牛潭组至宝塔组地层进行了详细的牙形类序列分析及对比。研究结果表明,自下而上可识别7个牙形类化石带,分别是Lenodus antivariabilis带、Lenodus variabilis带、Yangtzeplacognathus crassus带、Dzikodus tablepointensis带、Yangtzeplacognathus foliaceus带、Yangtzeplacognathus jianyeensis带及Hamarodus brevirameus带。剖面牙形类序列与华南中晚奥陶世综合序列基本一致,但缺失达瑞威尔晚期牙形类生物带(Eoplacognathus suecicus带至Yangtzeplacognathus protoramosus带)的完整记录,这一缺失在扬子台地多个剖面均有记录,可能指示达瑞威尔晚期存在的区域性海平面变化以及全球海洋古环境变化事件。关键词牙形类生物地层地层缺失牯牛潭组–宝塔组中–上奥陶统湖北地区     

陕西宁强—四川广元地区奥陶系“涧草沟组”和宝塔组牙形刺生物地层研究之新进展

开展陕西宁强范家沟、赵家坝和四川广元谭家沟3条剖面奥陶系牙形刺生物地层研究。在宁强地区"涧草沟组"发现了凯迪阶中部的Protopanderodus insculptus生物带,它相当于湖北宜昌地区上奥陶统临湘组之同名带;宝塔组由上而下可划分为Hamarodus brevirameus,Baltoniodus variabilis,Yangtzeplacognathus jianyeensis和Y.foliaceus等牙形刺带,属达瑞威尔阶上部至凯迪阶下部,可与湖北宜昌地区中-上奥陶统牯牛潭组顶部至宝塔组的同名带对比;广元地区宝塔组则见Baltoniodus variabilis和Yangtzeplacognathus jianyeensis生物带,属达瑞威尔阶顶部至桑比阶,可与湖北宜昌地区庙坡组或南京地区大田坝组的同名带对比。上述发现改变了先前的认识:陕西宁强地区宝塔组划分为Hamarodus europaeus和Eoplacognathus jianyeensis两牙形刺带,与湖北宜昌地区庙坡组和宝塔组地层相当;四川广元地区宝塔组归入Pygodus serra-Eoplacognathus foliaceus牙形刺带,与湖北宜昌地区牯牛潭组对比。邱洪荣(1989)建立的4个新种可能是一些老属种不同形态的组成分子,因此在本文属种描述部分把它们分别列入相应属种的同义名录。     

湖北宜昌真金和陈家河奥陶系牯牛潭组至宝塔组的牙形刺及其地层意义

根据采自湖北宜昌分乡陈家河和远安真金两剖面奥陶系牯牛潭组至宝塔组的牙形刺,将该段地层的牙形刺生物带自上而下综合划分为Hamarodus brevirameus,Amorphognathus superbus,Baltoniodus alobatus,Pygodus anserinus-Yangtzeplacognathus jianyeensis,Yangtzeplacognathus protoramosus,Yangtzeplacognathus foliaceus,Histiodella kristinae-Eoplacognathus suecicus,Dzikodus tablepointensis,Lenodus variabilis和Lenodus antivariabilis等牙形刺生物带。其中宝塔组之Hamarodus brevirameus,Amorphognathus superbus生物带及Baltoniodus alobatus生物带顶部归为凯迪阶(Katian),庙坡组Baltoniodus alobatus生物带大部和Pygodus anserinus-Yangtzeplacognathus jianyeensis生物带中上部归为桑比阶(Sandbian),而牯牛潭组的Yangtzeplacognathus protoramosus至Lenodus antivariabilis等牙形刺生物带和庙坡组Pygodus anserinus-Yangtzeplacognathus jianyeensis生物带下部则归入达瑞威尔阶(Darriwillian)。这一牙形刺带序列与宜昌黄花场和分乡剖面的牙形刺生物带相一致。     

鄂西奥陶系庙坡组海林檎Cheirocrinus penniger (Eichwald, 1842)的发现

鄂西地区中–晚奥陶世之交的庙坡组由数米厚的页岩夹少量薄层灰岩构成,沉积于扬子陆表海区凹陷带中,深度位于最大浪基面下,含有多样性、丰度高的化石组合。宜昌北部的陈家河剖面以及远安真金剖面庙坡组中–上部页岩属晚奥陶世桑比期Nemagraptus gracilis笔石带。笔者在该层位首次采获30余枚海林檎化石,经系统古生物学鉴定均为Cheirocrinus penniger (Eichwald, 1842)。远安真金剖面产出的C. penniger标本多以萼部区数个不同类型压扁的骨板形式保存, 1枚标本保存有完整的茎部;宜昌陈家河剖面的C. penniger标本均以单个萼部骨板形式保存。奥陶纪的Cheirocrinus广泛分布于多个板块,本文描记庙坡组的Cheirocrinus化石记录实例,扩展了该属在华南板块的时空分布,它可以从黔南三都早奥陶世斜坡相的同高组延续到晚奥陶世台内凹陷带的庙坡组。     

黔北上奥陶统宝塔组头足类多样性初探*

上奥陶统宝塔组(桑比阶–凯迪阶)是华南分布广泛的标志性地层单元, 以发育特殊的网纹状灰岩并产出大量头足类化石为特征。黔北桐梓县尧龙山地区的宝塔组厚度较大, 出露极好, 产出数量丰富、保存精美和特征典型的头足类化石。本文在研究区酒店垭、集镇采坑和天域极地3个宝塔组剖面采集到大量原域埋藏的头足类化石, 共8属11种及少量属种未定类型, 以直形壳或盘形壳、体管很小且适合远洋深水游泳的类型为主, 整体上符合扬子台地深水陆棚Sinoceras–Michelinoceras–Disoceras生物相特征。基于壳体扩大率、壳径与体管直径比、气室高与隔壁颈长比、隔壁下凹度、壳径与气室高比等参数变量的主成分分析表明, 在外部特征不显著的情况下, Sinoceras和Michelinoceras的直观差别是壳体扩大率, 而Michelinoceras属不同种的差异主要体现在气室高与隔壁颈长之比, 其他特征的区分度并不明显。层面上单位面积的化石定量统计显示, 尧龙山地区宝塔组中头足类的多样性较高, 并以Sinoceras和Michelinoceras具有极高的个体丰度为特征。扬子板块在奥陶纪桑比期–凯迪期之交处于高水位的陆表海环境, 栖息环境均一化和生态系统复杂化可能是促进该时期头足类动物的复苏和多样性演化的重要原因。     

湖北宜昌远安真金和界岭奥陶系牯牛潭组顶部的牙形刺及其地层意义（英文）

本文描述了采自湖北宜昌远安真金和界岭剖面奥陶系牯牛潭组顶部的牙形刺,它们有Baltoniodus prevariabilis,Drepanoistodus venustus,Panderodus gracilis,Periodon aculeatus,Protopanderodus calceatus,P.varicostatus?,Yangtzeplacognathus foliaceus和Y.protoramosus等,确认宜昌地区牯牛潭组顶部应归属于奥陶系达瑞威尔阶Pygodus serra带之Yangtzeplacognathus protoramosus亚带,并可与宜昌黄花场剖面和南京汤山外杆沟剖面同名组顶部及新疆柯坪羊吉坎剖面一间房组顶部的牙形刺带相对比。同时,它要比重庆城口大槽剖面牯牛潭组顶部的牙形刺带低3个带(Pygodus anserinus带至Baltoniodus variabilis带),这一事实也说明了牯牛潭组顶部在中国扬子地台的穿时性。     

安徽贵池早奥陶世晚期及中奥陶世的头足类

一、前言安徽贵池一带的奥陶系发育良好,含丰富的头足类化石。1984年夏,笔者等在贵池百安的方村剖面采获不少头足类化石,其中以下奥陶统的紫台组、牯牛潭组及中奥陶统的大田坝组、宝塔组所产的头足类最为丰富。这批化石经研究共有33种(其中10新种),分属15属8科。另有2未定属种。这些化石的发现增加了我国奥陶纪头足类的新材料,并对这个地区奥陶纪地层的划分以及与国内外的对比提供了可靠的依据。笔者在研究过程中,得到中国科学院南京     

江苏句容仑山附近奥陶纪鹦鹉螺

一、前言1984、1985年笔者与陈挺恩等两次对江苏句容仑山以及江宁汤山地区奥陶系剖面进行观察和研究.在仑山附近奥陶系剖面的红花园组、大湾组、牯牛潭组和宝塔组中,采到数量较多的鹦鹉螺化石.除红花园组化石制片尚有困难,暂时未着手研究;大湾组、牯牛潭组、宝塔组的化石,经研究共有13属、36种.其中1新属,ll新种,1比较种,2未定种.江苏句容县仑山,江宁县汤山附近的奥陶系研究较早.德国学者 Richthofen     

论中华震旦角石(Sinoceras chinense)的鉴别特征

晚奥陶世早期,随着海平面上升,鹦鹉螺逐渐成为海洋中的霸主,中华震旦角石[Sinoceras chinense(Foord)]开始繁盛。迄今为止Sinoceras chinense共记录有8个亚种,主要产出于晚奥陶世早期扬子区宝塔组网纹状灰岩之中。从前人的描述和图版来看,有些亚种与Sinoceras chinense模式标本差别较大,并且一些亚种并存于同一地区同一层位,这与生物学上种和亚种的定义相悖,需要重新厘定。因此,本文在归纳24篇前人文献中关于Sinoceras chinense的所有系统描述资料基础上,对迄今已发表的33个Sinoceras chinense标本选取6个主要壳体形态参数,运用聚类分析等多元统计分析手段,对这些标本的形态特征数据进行定量化分析,认为这些标本可以分为3个组合:Sinoceras chinense chinense组合、Sinoceras chinense eccentrica组合和非Sinoceras chinense组合,从而厘定了有关该种的分类问题,并明确Sinoceras chinense的鉴别特征,进而重新认识Sinoceras chinense的地理分布,认为Sinoceras chinense仅分布在我国华南、云南西部、新疆和西藏等地,华北地区和国外并没有确切的产出记录。     

扬子北缘钟祥地区奥陶系弗洛阶笔石动物群研究

湖北钟祥下奥陶统弗洛阶岩性以灰黑色页岩为主,夹粉砂岩及灰岩透镜体, 化石丰度和分异度高。通过系统的化石采集和鉴定, 于钟祥温峡口剖面识别出笔石7属31种, 大体上为扬子地台区的常见属种。自下而上分为Didymograptellus bifidus带、Corymbograptus deflexus带和Azygograptus suecicus带。该序列与宜昌?南漳地区一致, 但在笔石动物群组成面貌上差异较大,可能与钟祥地区在早奥陶世位于扬子台地边缘, 富集的营养物质和低氧的环境更利于笔石动物群的繁盛和笔石化石的保存有关。     

湘南地区晚奥陶世的江西笔石(Jiangxigraptus)*

本文对甘肃宝积山盆地中侏罗世植物群地质时代及植物区系进行了探讨, 共统计植物大化石 18 属 53 种, 主要包括银杏纲 10 属 24 种, 真蕨纲 5 属 22 种, 苏铁纲 1 属 3 种, 松柏纲 1 属 2 种, 楔叶纲 1 属 2 种。该植物群在组成特征上表现为银杏类、真蕨类植物繁盛, 其他植物类群相对贫乏。基于该植物群特征与国内其他植物群的对比, 提出其地质时代为中侏罗世 Aalenian 期–Bajocian 期。通过哈曼、欧式距离、罗杰斯–塔尼莫特和索卡尔–施尼斯 4 种方法对宝积山盆地及中国北方其余 11 个中侏罗世植物群进行系统聚类分析, 在此基础上进一步划分了中国北方植物区系。通过计算属、种相似性系数,探讨宝积山邻近植物区系间的亲缘关系, 结果表明宝积山植物区系与同属西北地区的华亭植物区系关系较为亲近。     

新疆准噶尔盆地白杨河中侏罗世植物化石新材料

系统描述了准噶尔盆地西北缘白杨河地区西山窑组植物化石15属25种,首次在新疆地区发现侏罗纪真蕨类重要分子Coniopteris hymenophylloides的原位孢子及其孢子囊群,并对Phoenicopsis ( Phoenicopsis) an-gustifolia的叶角质层进行了研究。白杨河地区西山窑组迄今已知植物化石24属39种,以富含蕨类、茨康类、银杏类及松柏类,少见苏铁类等为特征,属于较为典型的内陆型、温带或暖温带植物群;其时代为中侏罗世,且可能为中侏罗世早期。该植物群应属于欧亚大陆早、中侏罗世植物地理区系中的西伯利亚区,反映了温暖潮湿、具季节性变化、并有利成煤的环境。     

An Early Jurassic flora from the Clarence-Moreton Basin, Australia

A low-diversity Early Jurassic flora preserved in floodbasin siltstones of the Marburg Subgroup at Inverleigh Quarry in the Clarence-Moreton Basin, eastern Australia, is dominated by Allocladus helgei Jansson sp. nov., a conifer with denticulate leaves tentatively attributed to Araucariaceae. The assemblage also includes Rintoulia variabilis and Caytoniales, (Caytonia cucullata McLoughlin sp. nov. and cf. Sagenopteris nilssoniana), reinforcing the wide distribution of this order in Early to Middle Jurassic floras of Gondwana. Ferns (Cladophlebis and Sphenopteris species) and isoetalean lycophytes (Isoetites sp.) constitute the herbaceous elements of the flora. The palynoflora is dominated by cheirolepidiacean (Classopollis) pollen and is attributable to the upper part of the Corollina (= Classopollis) torosa Zone of late Pliensbachian–early Toarcian age (180–185 Ma). The Inverleigh flora represents one of the few Australian assemblages dated between the major phases of floristic turnover at the end of the Triassic and the Toarcian. Sedimentological characteristics, cuticular features of the conifer leaves and the abundance of free-sporing plants indicate a relatively humid palaeoclimate for the Clarence-Moreton Basin Early Jurassic.     

Phytogeography of the bryophyte floras of oak forests and páramo of the Cordillera de Talamanca, Costa Rica

Aim Central America is a biogeographically interesting area because of its location between the rich and very different biota of North and South America. We aim to assess phytogeographical patterns in the bryophyte floras of oak forests and páramo of the Cordillera de Talamanca, Costa Rica. Location Tropical America, in particular the montane area of Cordillera de Talamanca, Costa Rica. Methods The analysis is based on a new critical inventory of the montane bryophyte flora of Cordillera de Talamanca. All species were assigned to phytogeographical elements on the basis of their currently known distribution. Absolute and percentage similarities were employed to evaluate floristic affinities. Results A total of 401 species [191 hepatics (liverworts), one hornwort, 209 mosses] are recorded; of these, 251 species (128 hepatics, one hornwort, 122 mosses) occur in oak forests. Ninety‐three per cent of all oak forest species are tropical in distribution, the remaining 7% are temperate (4%) and cosmopolitan (3%) species. The neotropical element includes almost 74% of the species, the wide tropical element (pantropical, amphi‐atlantic, amphi‐pacific) only 19%. A significant part of the neotropical species from oak forests are species with tropical Andean‐centred ranges (27%). As compared with bryophyte species, vascular plant genera in the study region are represented by fewer neotropical, more temperate and more amphi‐pacific taxa. Bryophyte floras of different microhabitats within the oak forest and epiphytic bryophyte floras on Quercus copeyensis in primary, early secondary and late secondary oak forest show a similar phytogeographical make‐up to the total oak forest bryophyte flora. Comparison of oak forest and páramo reveals a greater affinity of the páramo bryophyte flora to temperate regions and the great importance of the páramo element in páramo. Surprisingly, oak forests have more Central American endemics than páramo. Main conclusions (1) Providing first insights into the phytogeographical patterns of the bryophyte flora of oak forests and páramo, we are able to confirm general phytogeographical trends recorded from vascular plant genera of the study area although the latter were more rich in temperate taxa. (2) Andean‐centred species are a conspicuous element in the bryophyte flora of Cordillera de Talamanca, reflecting the close historical connection between the montane bryophyte floras of Costa Rica and South America. (3) High percentages of Central American endemics in the bryophyte flora of the oak forests suggest the importance of climatic changes associated with Pleistocene glaciations for allopatric speciation.     

Middle Jurassic Flora from Shouthern Margin of Eerduosi Basin

This paper deals with some Middle Jurassic fossil plants from southern margin of the Eerduosi Basin. There are 36 species in 13 genus in this flora, i.e. Equisetum multi- dentatus Oishi, E. sp., Coniopteris hymenophylloides (Brongn.) Seward, C. bella Harris, C, burejensis (Zalessky) Seward, C. szeiana Choweet Yeh, C. tatungensis Sze, C. spp., Todites williamsoni (Brongn.) Seward, T. cf. williamsoni (Brongn.) Seward, Cladophle- bis asiatica Chow et Yeh, C. argutula (Heer) Fontaine, C. hsiehiana Sze, C. ( Gleichenites) takeyamae Oishi et Takahasi, C. cf. gracils Sze C. triangularis Oishi, C. grabauiana P’an C. cf punctata (Thoms) Chow et Yeh, C. spp., Baiera cf. concinna (Heer) Kawasaki, B. furcata (L. et H.) Braun, B. sp., Sphenobaiera longifolia (Pomel) Florin, Ginkgo obrutchewi Seward, G. sp., Phoenicopsis angustifolia Heer P. manchurica (Yo- koyama) Yabe, p. sp., Czekanowskia rigida Heer, C. sp., Pterphyllum sp., Elatocladus cf manchurica (Yokoyama) Yabe, E. sp., Podozamites lanceolatus (L. et. H.) Braun P. sp., Carpolithus sp. etc. The dominant groups of this flora are Filicopsida and Ginkgopsida. So it could be compared with the flora of Yanan Formation from inner Earduosi Basin and other floras from Northeast, Northwest and China, such as the floras of Dutung, Mentougon and Dongsheng etc. This flora is different from the Late Triassic flora and the Early Jurassic flora of North China. It belongs to the Coniopteris-Phoenicopsis flo- ra, which was assigned to Early-Middle Jurassic flora by Prof. Sze. But after studying, the authers believe that this flora should belong to Middle Jurassic rather than Early Jurassic. The climate was warm and damp in that time in North China.     

Observations on the formation of Chinese desert floras

1. Based upon the analyses in the floristic elements of the three genera (Suaeda, Salsola and Zygophyllum) in different regions we can see that the genesis of our desert floras in these regions is very much diversified. The flora of Songaria is similar to that of the Middle Asia, while the Hosi Corridor seems to be a transitional area very close to Alashan and also related to the Tarim Basin in floristic elements. Thus, we may classify the desert floras into three parts: the flora of Songaria, of Alashan including the Hosi Corridor and of the Tarim Basin including the Tsaidam Basin. The ages and approaches in their formation are different. 2. There are plenty species but no or rare endemics in Songaria. In spring there are a number of ephemeral plants. The variation of aspect is evident. The vegetation cover is abundant. The floristic elements are developed from the flora of Middle Asia and it was formed in Quaternary period. 3. The floristic elements of the Tarim Basin are poor, but there are not few endemics and the distribution of the endemics is much limited. They are of the characteristics of relic species. Therefore it was formed in the Tertiary period and developed in Quaternary period. The elements are related to the Mediterranean flora. 4. There are a large number of endemics and many endemic monotypie genera in Alashan. They represent the flora formed in Tertiary period. Although it is of a special style, it relates both to the Middle-Asian and the Mediterranean flora. 5. The historic causes for the formation of the different floras lie chiefly on: (1) The rise of the Tibetan plateau and mountains strongly changed the climatic and edaphic conditions and in the long course of evolution some species survived or even developed, while the others deteriorated or even died out from the flora. (2) Because the circumstances of transgression or regression of the Tethys were different in these regions. (3) The mountain-making movement, the transgression and regression and the fluence of glaciation, all the mutation of these associated factors modified the climatic zonation and then the plant species changes followed, new species formed and migration of floristie elements occurred. (4) Songaria is the nearest region to the then Sibirian glacier, so the frozen injury to the flora might be the greatest. (5) In the Glacial period the descension of snow line in Songaria was greater than that of the Tarim Basin, so the frozen injurymight be greater.     

A phytogeographical characterization of the vine flora of the Sonoran and Chihuahuan deserts

Aim This study presents a phytogeographical characterization of the vine flora of two lower North American desert regions as a biogeographical framework for further ecological inquiry into desert vines. Location The phytogeography of the vine flora of the Sonoran and Chihuahuan Deserts was c haracterized based on 263 known species. Methods Checklists of the vines of each desert were developed. Represented genera were then grouped into 10 phytogeographical elements based on worldwide distribution patterns. To compare the floristic composition of the desert floras, an index of species similarity was calculated. Results About a third more species of vines occur in the Sonoran Desert than in the Chihuahuan Desert. Based on the analysis, cosmopolitan genera are the only group more numerous in absolute terms in the Chihuahuan Desert than in the Sonoran Desert. Tropical elements are represented in about the same proportion in each desert as the number of species, however, nearly twice as many pantropical and neotropical genera are represented in the Sonoran Desert as in the Chihuahuan Desert. Proportionately, more genera of temperate elements occur in the Chihuahuan Desert than in the Sonoran desert, although the absolute number of genera is slightly higher in the latter. Main conclusions As these deserts are relatively recent ecological formations and as vines evolved in forest ecosystems, the composition of the desert vine floras is the result of the interaction between historical vegetation types, their constituent taxa and climatic and geological history. The main differences in the vining floras of the present‐day Sonoran and Chihuahuan Deserts appear to be the result of greater historical influence in the Sonoran Desert of (1) tropical vegetation types and (2) the emergence of the Gulf of California. The Chihuahuan Desert vine flora seems to be the result of (1) a more pronounced historical temperate vegetation, (2) the lack of an important isolating event, such as the creation of the Baja California peninsula, and (3) a cooler climate with shorter growing seasons.     

The taxonomy and ecology of the vascular plant flora of Singapore: a statistical analysis

A total of 2277 species of vascular plant in 868 genera and 158 families are recorded as native to the Republic of Singapore. This flora consists of 174 pteridophytes (7.6% of the total), eight gymnosperms (0.4%), 590 monocots (25.9%) and 1505 dicots (66.1%).Median values of species per family, genera per family and species per genus are 5, 2 and 1, respectively. The most abundant 'families' and genera in terms of species are Orchidaceae (196 spp.), Pteridophyta (174 spp.), Rubiaceae (140 spp.), Euphorbiaceae (116 spp.), Gramineae (100 spp.) and Eugenia (46 spp.), Ficus (43 spp.), Dendrobium (33 spp.), Bulbophyllum (29 spp.) and Cyperus (29 spp.). Trees constitute 37.2% of the flora, shrubs 9.8%, herbs 25.9%, climbers 14.0% and epiphytes 13%.Euphorbiaceae and Rubiaceae dominate the woody flora; pteridophytes, Gramineae and Cyperaceae the herbs; Rubiaceae, Palmae and Annonaceae the climbers and Orchidaceae and pteridophytes the epiphyteS. Species confined to coastal habitats comprise 11.1% of the total flora; 73.5% are lowland forest species and the remaining 15.5% are non-tree species of open and frequently-disturbed sites. The vascular plant of flora of Bukit Timah Nature Reserve, the largest remaining patch of primary rain forest in Singapore, is also analysed. Its flora is similar to that of the forest component of the complete Singapore flora. The Singapore flora, in comparison with other tropical floras, is highly diverse, but appears depauperate in epiphytes and climbers.     

中国主要侏罗纪植物群相似性系数的初步研究

选取属级分类单元作为统计变量, Dice系数作为统计公式, 对中国早中侏罗世较具代表性的18个植物群进行了相似性的聚类分析。研究结果显示出的区系划分格局在一定程度上支持了前人对植物群时代和性质的判断, 同时提出香溪组、万宝组所含植物群的特殊组合与时空因素值得深入研究; 中国北方早侏罗世植物群、北方中侏罗世植物群以及南方早中侏罗世植物群三者之间的差异、联系及范围界限 需要进一步深入研究。本研究是对古植物学区系研究的有益探索, 为中国侏罗纪植物群的大区域系统研究提供了数据基础和参考依据。     

中国主要侏罗纪植物群相似性系数的初步研究

选取属级分类单元作为统计变量,Dice系数作为统计公式,对中国早中侏罗世较具代表性的18个植物群进行了相似性的聚类分析.研究结果显示出的区系划分格局在一定程度上支持了前人对植物群时代和性质的判断,同时提出香溪组、万宝组所含植物群的特殊组合与时空因素值得深入研究;中国北方早侏罗世植物群、北方中侏罗世植物群以及南方早中侏罗世植物群三者之间的差异、联系及范围界限需要进一步深入研究.本研究是对古植物学区系研究的有益探索,为中国侏罗纪植物群的大区域系统研究提供了数据基础和参考依据.