山地次生林破碎化对喜鹊繁殖功效的影响

于1999－2000年春夏两季,在左家自然保护区及土门岭地区研究了破碎化山地次生林斑块面积对喜鹊（Pica pica）繁殖功效的影响。运用GPS定位系统确定了12块大（＞20hm^2,n=4）、中（10－20hm^2,n=hm^2）对照样地（每块面积20hm^2）。观测了各斑块及对照样地中喜鹊产第1枚卵的时间、窝卵数、平均卵重、出雏量及雏鸟出飞量等生态指标,结果表明,（1）斑块面积对喜鹊的繁殖功效存在着显著的影响;（2）喜鹊在对照样地和大面积斑块中的平均产卵时间早于小斑块中的11.62d;（3）对照样地和大面积斑块内的窝卵数略高于中、小面积块内,但差异不显著;（4）各斑块及对照样地中的平均卵重无显著差异,小面积斑块中的出雏率和雏鸟出飞率均最低（51.7%和42.88%）,大面积斑块中的出雏率最高（72.12%）,对照样地略次之（71.93%）,对照样地中的雏鸟出飞率最高（65.45%）,大面积斑块次之（62.71%）;（5）喜鹊在小面积次生林斑块中繁殖功效较低的主要原因是巢损失率较高。     

温带次生林破碎化对白眉姬鹟繁殖的影响

于2005～2006年鸟类繁殖季节,通过悬挂人工巢箱,探讨了次生林破碎化对白眉姬鹟Ficedula zanthopygia繁殖的影响.结果 表明:次生林破碎化对白眉姬鹟繁殖产生很大的影响,斑块面积、形状指数、隔离度均在不同程度上影响着白眉姬鹟的繁殖参数,破碎化指数不同的斑块间白眉姬鹟的窝卵数、孵化率和出飞率差异均不显著(P＞0.05);卵重变异很大,差异显著(P＜0.05);卵体积变动更大,差异达到极显著水平(P＜0.01).回归分析表明,斑块破碎化指数与白眉姬鹟繁殖参数间均存在一元二次方程关系:当窝卵数达到最大值时,卵质量降低,出雏率达到最低,雏鸟存活率反而增大,出飞率达到最大,从而提高繁殖功效;当窝卵数达到最小值时,卵质量最高,出雏率增大,出飞率也相应的增高,从而使繁殖功效增大.白眉姬鹟存在一个最优窝卵数,验证了Lack的窝卵数假说.另外,白眉姬鹟通过对自身繁殖参数的调节来适应生境破碎化,尽量提高自身的繁殖功效,从而弥补自身种群.     

人工巢箱中大山雀和褐头山雀的繁殖比较

2009年3-7月,在北京小龙门森林公园悬挂人工巢箱,对大山雀(Parus major)和褐头山雀(P.montanus;)的繁殖进行比较研究.野外共悬挂100个巢箱,其中19巢(19.0%)被大山雀、11巢(11.0%)被褐头山雀进驻,总利用率为30.0%.同域繁殖的大山雀和褐头山雀在窝卵数、孵化期上有极显著差异(P＜0.01),其中大山雀的窝卵数(8.21枚±0.25枚,n=19)极显著大于褐头山雀(6.18枚±0.23枚,n=11),而褐头山雀的孵化期(14.22d±0.44d,n=10)极显著长于大山雀(13.17 d±O.83 d,n=12).两者在卵重、卵大小、出雏数、雏鸟出飞数上无显著差异(P＞0.05).大山雀的繁殖成功率(27.0%)和营巢成效(63.2%)均显著低于褐头山雀(54.5%和100%)(P＜0.01).表明大山雀可能采取高产卵数、低存活率的繁殖对策,而褐头山雀的繁殖策略则可能为低产卵数、高存活率.     

筑巢投入与巢材组成对大山雀繁殖产出的影响

鸟类在筑巢期投入能量的多少会影响其繁殖成效,巢材组成的差异也会影响其最终繁殖产出。本文以大山雀(Parus major)为研究对象,通过探究巢重及巢材组成对其繁殖的影响,以揭示筑巢投入和巢材对繁殖产出的影响。对2015年收集到的35巢进行了分析。结果表明:巢材总重与出飞数、出飞成功率及繁殖成功率呈极显著的负相关;羽毛重及其所占总重的比例与窝卵数、出雏数及出飞数呈显著的正相关,与产首枚卵的日期呈显著的负相关;植物茎重及其所占总重的比例与窝卵数及出雏数呈显著的正相关。本研究表明,大山雀亲鸟筑巢消耗的能量越多,子代出飞情况反而越差;同时,巢内保温性和支撑性巢材越多,繁殖产出越大。     

两种雀形目鸟类孵化行为对子代质量的影响

对高寒草甸地面营巢的小云雀 (Alaudagulgula)和灌丛筑巢的黄嘴朱顶雀 (Acanthisflavirostris)繁殖方式的研究表明 :①小云雀孵化异步性较弱 (1d) ;平均卵重与产卵顺序不显著相关 ,但最后 1枚卵最重 ;出壳顺序与雏鸟生长率极显著正相关 (P <0 0 1) ;出壳顺序对幼鸟的雏期和离巢体重无显著影响 (P >0 0 5 )。②黄嘴朱顶雀孵化异步性较强 (2～ 3d) ;平均卵重与产卵顺序显著正相关 (P <0 0 5 ) ,最后 1枚卵最重 ,第 1枚次之 ;出壳顺序对雏鸟生长率、雏期和离巢体重均无显著影响 (P >0 0 5 )。结果分析表明 ,2种雀形目鸟类都选择了异步孵化和加强窝雏数对策的繁殖模式。     

太湖地区夜鹭配对年龄及繁殖效果

为了解夜鹭同龄配对及其与窝卵数、窝雏数、雏鸟成活率的关系,2001年4-6月对无锡太湖地区夜鹭（Nycticorax nycticorax）的年龄配对及其繁殖作了观察。在方便观察的206个样巢中,二龄（2Y）、三龄（3Y）鸟参与配对繁殖的达35.4％：亲鸟“2Y 4Y^ “的2对,“2Y 2Y“的39对,“3Y 3Y“的34对。“4Y^ 4Y^ 的131对（含6对弃巢鸟）。在200个成功育雏的巢中,不同年龄亲鸟所产卵的体积和窝卵数无差异,但卵重、窝雏数、孵化率及成活率有随亲鸟年龄增加的趋势。     

厦门鸡屿岛白鹭几种繁殖活动的观察

于 2 0 0 1年 3～ 7月采用颜色标记法 ,记录厦门白鹭保护区内鸡屿岛白鹭 (Egrettagarzetta)卵产出和孵出的顺序和时间 ,并称重卵和雏鸟。有 91 4 %的窝卵数为 4～ 5枚 ,产卵期 7 0± 1 9d ,出壳期 4 2± 1 4d ;卵孵化时间与产卵顺序显著负相关 ;不同产出顺序卵重无差异 ;不同孵出顺序雏鸟早期 (≤ 5d)发育无差异 ,之后差异显著 ,第 4出壳的雏鸟发育水平和成活率较低 ,而第 5出壳的雏鸟最低 ;亲鸟在产卵期的孵化是非连续的 ,而产卵结束后相对连续。如此 ,可以调节孵化时间 ,进而调控异步孵化的程度 ,使雏鸟在生长阶段形成适当的等级差别 ,以获得最大的繁殖收益。     

两种雀形目鸟类的窝雏数处理实验:检验Lack假说

于1997—1999年在位于青海省北部的中国科学院海北高寒草甸生态系统定位站进行。选择地面筑巢的小云雀(Alauda gulgula)和灌丛筑巢的黄嘴朱顶雀(Acanthis flavirostris)为代表进行窝雏数处理实验。根据Lack假说的预报检验(1)常见窝卵数是否是最大生产力窝卵数;(2)窝雏数处理对雏鸟质量和亲鸟投入是否产生影响;(3)两种鸟的响应方式是否相同。其结果如下：①小云雀和黄嘴朱顶雀的常见窝卵数分别是3和5枚。年间变化不明显,用幼鸟出飞率作为生产力,两种鸟的扩增窝幼鸟出飞率下降,常见窝卵数(分布频率最高)等同于最大生产力窝卵数;②小云雀的幼鸟的生长参数不随窝雏数的改变而变化．而黄嘴朱顶雀有明显变化．说明窝雏数处理对后者幼鸟质量有明显影响。③用递食率作为亲鸟投资指标,小云雀亲鸟的递食率随窝雏数的增加而增加,但雏期不变;而黄嘴朱顶雀递食率不变,但雏期延长。④扩增窝雏数后,两种亲鸟表现出不同的响应方式,小云雀表现为提高单位时间递食次数,而黄嘴朱顶雀延长育幼时间。这两种方式不是通过影响雏鸟质量就是通过影响亲鸟存活率来降低子代和亲代的适合度。结果支持了自然选择将窝卵数调节到亲鸟能喂活最大数量子代的限度。即常见窝卵数就是最大生产力窝卵数的Lack假说。     

江西吉安乌鸫的繁殖生态研究

2009～2010年对江西省吉安地区乌鸫Turdus merula的繁殖进行了调查研究,结果表明:当地乌鸫的营巢时间在3月9日至16日,产卵时间是3月18日至26日,平均窝卵数5.14(4～6)枚(n=14),平均卵大小29.71 mm×21.16 mm,平均卵重6.63 g(n=71).孵卵和暖巢主要由雌鸟承担,但雄鸟也有暖巢行为,孵化率为65.83%,育雏期13～15 d,离巢率高达100%.与高海拔的乌鸫相比,当地乌鸫产大窝小卵,这一特征保证了大窝雏数和高离巢率,繁殖对策属于r-选择.     

骨顶鸡的繁殖行为

2008和2009年4-9月,对黑龙江省安邦河自然保护区以及龙凤湿地骨顶鸡繁殖种群进行了观察.结果表明:骨顶鸡繁殖季节会建造3种巢结构:炫耀台、产卵巢和育雏巢,雌雄共同参与筑巢及领域防卫活动;巢高和巢深随窝卵数增多而呈上升趋势;平均窝卵数为8.77±0.24(n=75)枚,窝卵数随繁殖时间推迟而呈减少趋势;骨顶鸡雌雄共同参与孵化,雌性平均每次孵化时间高于雄性;雌雄共同参与喂雏,雏鸟至亚体阶段才完全独立,之前需亲鸟饲喂;骨顶鸡是杂食性鸟类,食物以植物为主,但也摄取动物性食物.     

The effect of old age on the reproduction of Great Tits Parus major and Blue Tits P. caeruleus

The results from two data sets show that in Great Tits Parus major and Blue Tits P. caeruleus reproduction is influenced by age. In both species, laying dates become earlier while clutch- and brood-size increase between the ages of 1 and 2, but there is no change in nesting success or post-fledging juvenile survival. Great Tits aged 5 or older are 'old' in that laying starts later and nesting success, brood-size and post-fledging juvenile survival decrease. Blue Tits become 'old' 1 year earlier than Great Tits: females aged 4 or older lay later, have a lower nesting success and smaller brood-size, and their young show a decreased post-fledging survival. The effect on laying date, in the Blue Tit, becomes apparent only for females aged 6 and older. It is concluded that ageing in small passerines is to be expected more generally, but that no extrapolations can be made as to the timing of the effect from one species to another.     

Effect of age,haemosporidian infection and body condition on pair composition and reproductive success in Great Tits Parus major

Several factors shape lifetime reproductive success, including genetic background, body condition, environmental conditions and ecological interactions such as parasitism. Adults often show higher reproductive success than their young conspecifics, especially in long-lived bird species, and this may be explained by the cumulative effects of an increase in reproductive experience and the selection of high-quality individuals from one year to the next. To test whether this pattern also exists in short-lived bird species, we used 13 years of monitoring data from two Great Tit Parus major populations. The effects of male and female age on several reproductive parameters were analysed in 419 pairs of Great Tits, while accounting for body condition and infection by haemosporidian parasites. Reproductive success was mainly affected by the age-class of males. Pairs containing a sub-adult male fledged one-third fewer chicks than pairs containing an adult male. The difference was not caused by variation in male fertility but could have been caused by better parental care provided by adult birds. In addition to lower reproductive success, first-year males also had reduced access to mating compared with adult males, suggesting an avoidance of sub-adult males by females. Nestling body condition was positively correlated with parental body condition, and the body condition of male and female members of breeding pairs was positively correlated. Finally, the number of fledged chicks was mainly affected by the infection status of males. These results temper our previously published results showing an effect of infection on Great Tit reproduction regardless of their sex. In our previous study, and as in most cases, the status of the partner was not taken into account and we show here that this is essential because it can lead to a biased interpretation of the results.     

Distribution and reproductive success of the Pied Flycatcher Ficedula hypoleuca in relation to forest patch size and vegetation characteristics; the effect of scale

We studied habitat choice, distribution and reproductive success in the Pied Flycatcher Ficedula hypoleuca in relation to the spatial structure of a fragmented forest area. Two hierarchical levels of forest habitat structure were used: (1) forest patch size (macrohabitat) and (2) vegetative structure within forest patches (microhabitat). In spring, both males and females settled preferentially in large and medium-sized forest stands (>1 ha) where breeding density was also higher than in small stands (<1 ha). Stands <5 ha were occupied later, and the proportion of unpaired males was higher there than in large stands (>5 ha). The known age distributions of breeding birds and breeding success were independent of forest patch size. Nest predation rate was not associated with stand size or nest distance from the forest edge. The preference of breeding birds for large forest patches was presumably related to the higher amount of resources (nest sites, mates, food) a large patch can offer for reproduction compared with a small patch. At the microhabitat level, territories of old males were characterized by relatively more deciduous trees that contained more invertebrate food than coniferous tree-dominated territories of yearling males. The reproductive success of old males, as measured by the number of fledged young per male, was higher than that of yearling males. This suggests that the larger body size and blacker plumage of old males possibly contributed to male dominance and that old males excluded younger males from preferred habitats through territorial behaviour. Our results suggest that habitat choice of the Pied Flycatcher was affected by both micro- and macrohabitat. On the basis of settlement pattern, density and reproductive success, the distribution of Pied Flycatchers across forest stands of different sizes followed the ideal-free distribution model, whereas at the microhabitat level, age-related unequal distribution of males followed the ideal-despotic model of Fretwell and Lucas.     

Higher climate warming sensitivity of Siberian larch in small than large forest islands in the fragmented Mongolian forest steppe

Forest fragmentation has been found to affect biodiversity and ecosystem functioning in multiple ways. We asked whether forest size and isolation in fragmented woodlands influences the climate warming sensitivity of tree growth in the southern boreal forest of the Mongolian Larix sibirica forest steppe, a naturally fragmented woodland embedded in grassland, which is highly affected by warming, drought, and increasing anthropogenic forest destruction in recent time. We examined the influence of stand size and stand isolation on the growth performance of larch in forests of four different size classes located in a woodland‐dominated forest‐steppe area and small forest patches in a grassland‐dominated area. We found increasing climate sensitivity and decreasing first‐order autocorrelation of annual stemwood increment with decreasing stand size. Stemwood increment increased with previous year's June and August precipitation in the three smallest forest size classes, but not in the largest forests. In the grassland‐dominated area, the tree growth dependence on summer rainfall was highest. Missing ring frequency has strongly increased since the 1970s in small, but not in large forests. In the grassland‐dominated area, the increase was much greater than in the forest‐dominated landscape. Forest regeneration decreased with decreasing stand size and was scarce or absent in the smallest forests. Our results suggest that the larch trees in small and isolated forest patches are far more susceptible to climate warming than in large continuous forests pointing to a grim future for the forests in this strongly warming region of the boreal forest that is also under high land use pressure.     

温带次生林的岛屿化对鸟类物种多样性及密度的影响

由于自然事件的影响和人类活动的干扰,越来越多的大片森林破碎成彼此孤立、面积不一的森林岛屿,这种变化无疑会对某些动物的分布模式及行为特征产生影响。于2000和2001年的春夏季,在吉林省左家自然保护区及土门岭地区,采用点样法对18块森林岛屿(面积范围:4.3–76.9hm2)中的鸟类物种多样性及密度进行了调查。主要目的是检测森林岛屿的面积效应是否对鸟类物种多样性及密度产生影响,同时分析经典的岛屿生物地理理论是否可以解释破碎化后的森林岛屿面积与物种的关系。结果表明,鸟类物种多样性在年间没有显著变化,但鸟类的密度在不同年间变化较大。不同面积森林岛屿中的鸟类物种多样性有所差异,所包含的鸟类物种数从12种到43种不等。尽管有些面积较大的斑块所包含的物种数较少,但鸟类物种数的总体趋势是随着斑块面积的增大而增多。不同鸟类对森林岛屿面积的反应并不相同,灰椋鸟(Sturnuscineraceus)、红尾伯劳(Laniuscristatus)、灰头鹀(Embrizaspodocephala)等在面积较小的斑块中密度较大,而山鹡鸰(Dendronanthusindicus)、树鹨(Anthushodg-soni)、灰背鸫(Turdushortulorum)等几乎不分布于小面积斑块之中。森林岛屿中鸟类物种随着面积变化的变异方式符合经典的岛屿生物地理理论的基本模式,但Z值和C值差异较大     

Fortpflanzungsbiologie der KohlmeiseParus major auf kleinen Fl?chen: Vergleich zwischen einheimischen und exotischen Baumbest?nden

Zusammenfassung Im Exotenwald von Weinheim wurden brutbiologische Untersuchungen an Kohlmeisen durchgeführt. Ziel der Untersuchungen war es, am Beispiel eines Höhlenbrüters exemplarisch festzustellen, ob nichteinheimische Vegetationselemente die Reproduktionsbedingungen beeinflussen können. Brutpaardichte und Reproduktionserfolg, sowie Körpermasse-Entwicklung von Kohlmeisennestlingen, Nahrungsangebot und Nahrungssuchverhalten wurden hierzu in exotischen und einheimisch bewaldeten UF vergleichend analysiert. Brutbiologische Parameter zwischen Brutpaaren in exotischen und einheimisch bewaldeten UF unterschieden sich nicht. In einheimisch bewaldeten Gebieten siedelten mehr Kohlmeisen und die flächenbezogene Reproduktionsrate war höher als in exotischer Bewaldung. In einheimischen Beständen mit größerem Raupenangebot wuchsen schwerere Jungvögel heran als in exotischen Beständen mit geringerer Raupendichte. Die Nahrungssuche fütternder Altvögel richtet sich bevorzugt auf Laub- und Obstbaumarten der einheimischen Vegetation. Exotische Nadel- und Zypressenvegetation wird als wenig ertragreiche Ressource gemieden. Die vorliegenden Ergebnisse zeigen, welch hohen Stellenwert einheimische Baumarten in unseren Forsten innehaben. Es ergibt sich als Empfehlung aus ornithologischer Sicht bei zukünftigen Aufforstungen von exotischen Baumarten den Anteil von Nadel- und Zypressenhölzern zu Gunsten von Laubbaumarten zu mindern und diese noch mit einheimischen Baumarten zu durchmischen.

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The reproductive biology of the Great TitParus major in small patches: exotic versus native tree species

In SW-Germany (Weinheim, 54.89 N; 34.76 E) some aspects of breeding biology of Great Tit were studied in forest patches with native versus exotic trees. Density of breeding pairs, breeding success, development of nestlings, food supply and foraging behaviour of feeding adults were compared between five patches with native and ten patches with exotic trees. Clutch size, hatching and fledging rates did not differ between the two vegetation types. Breeding densities as well as the patch size related production of hatched and fledged young were higher in native than in exotic forest patches. Caterpillar density was higher in native vegetation and the body mass of nestlings in such patches were clearly heavier than those of birds raised in exotic vegetation. Feeding adults in all vegetation types left the breeding patches to search for food but birds from exotic patches did this to a higher degree. Adults, especially those from native vegetation avoided exotic trees while foraging. Broad leafed exotic tree species were sometimes searched only by birds breeding in such vegetation. If exotic trees are favoured by afforestation measurements, a reduction of coniferous and mixing with native trees should be preferred to improve these habitats for birds.

     

Distribution and size of capercaillie leks in relation to old forest fragmentation

Summary Distribution and size of 38 capercaillie Tetrao urogallus leks were related to amount and configuration of old forest patches in two south-east Norwegian coniferous forests. The smallest occupied patch was 48 ha containing a solitary displaying cock. All patches larger than 1 km² contained leks. Number of cocks per lek increased with increasing patch size. Number of leks per patch increased in a step-wise manner with one lek added for each 2.5–3 km² increase in patch size. In large patches there was one lek per 3–5 km² old forest, and density of lekking cocks was 2–2.5 per km². In small patches density of cocks varied considerably. Density of cocks was not related to patch isolation or patch shape. However, among leks surrounded by 50–60% old forest within a 1 km radius, number of cocks increased with increasing old forest fine-graininess. We argue that when old forests cover more than 50%, a fine-grained mosaic may support higher densities of lekking cocks than a coarse-grained mosaic. Conversely, when old forests cover less than 50%, a fine-grained mosaic is unfavourable, because each old forest patch becomes too small and isolated. Finally, we present a predictive model of how old forest fragmentation influences density of leks, number of cocks per lek, and total density of cocks.     

Trait-specific response of ground beetles (Coleoptera: Carabidae) to forest fragmentation in the temperate region in Korea

The objective of this study was to examine the trait-specific response of ground beetles in terms of abundance, species richness, and composition for habitat fragmentation in temperate forests. In addition, we examined how different ecological groups and species of ground beetles responded to environmental variables. During middle May to early November in 2013, ground beetles were sampled using pitfall traps in 27 sites (18 patches and 9 continuous forests) in central Korea. A total of 51 species were identified from 17,845 ground beetles in the 27 study sites. Continuous forests had generally higher abundance and species richness of ground beetles than forest patches. Canonical correspondence analysis for ground beetles was significant, and the proportion of variance explained by environmental variables, such as patch size, elevation, organic matter (C), and soil moisture and pH, was 43.61%. In addition, species composition of ground beetles in continuous forests was grouped distinctively away from those in medium and small-sized patches. In conclusion, both small (1.1–9.6 ha) and medium patches (12.8–51.2 ha) were failed to preserve similar ground beetle biodiversity compared to continuous forests. However, our study revealed that medium-sized forest patches clearly had higher conservation value for forest specialist ground beetles than small-sized forest patches irrespective of forest type.     

Effects of Patch Size and Floral Herbivory on Seed Set in Trichogoniopsis adenantha (Asteraceae) in Southeastern Brazil1

The effects of patch size and floral herbivory on reproductive success of Trichogoniopsis adenantha (Asteraceae) were studied in a semideciduous forest in southeastern Brazil. Plants in large patches and capitula not attacked by endophagous insects had higher seed set. The consequences of living in large or small patches for plant reproductive success are discussed.     

利用天然树洞繁殖的五种鸟的巢位特征及繁殖成功率

对吉林省左家自然保护区次生阔叶林中的大山雀 ( Parusmajor)、沼泽山雀 ( Paruspalustris)、普通跓( Sitta europacea)、白眉姬? ( Ficedula zanthopygia)和灰椋鸟 ( Sturnus cineraceus) 5种利用天然树洞繁殖的次级洞巢鸟进行了巢位选择和繁殖成功率研究。本研究中共发现 1 41巢。五种鸟对树洞类型的选择存在种间差异 ,普通跓不利用裂洞 ,沼泽山雀不利用啄洞 ,其它 3种鸟对 3种洞均有利用 ,但有一定的倾向性。对 5种鸟 9个巢位变量的比较中 ,只有洞口方向差异不显著 ( p >0 .0 5 ) ,其它 8个变量均差异显著 ( p<0 .0 5 ) ,该结果说明 5种次级洞巢鸟对巢位的选择具有其各自的需求。洞口横径、洞口纵径、洞处树直径、洞内径、巢距地高是巢位选择重要变量 ,它们决定不同种类对树洞的利用。巢损失多数出现在产卵之前和孵化阶段 ,44个繁殖失败的巢中有 35个在这两个阶段损失。大山雀的巢成功率最低 ,灰椋鸟的巢成功率最高。 5种鸟的孵化率都超过 90 %。人为破坏和动物捕食是繁殖失败的主要原因 ,占总数的 61 .4%。洞巢鸟巢位选择中的重要变量影响繁殖成功。普通跓繁殖是否成功受洞口横径和巢高影响 ,沼泽山雀受洞口纵径、树胸径和洞内径影响 ,大山雀受洞口横径、巢高和洞内径影响 ,灰椋鸟受洞内径和洞深影