中国莎草科植物新资料

报道了产于西藏南部的1 个中国莎草科分布新记录种--阔鳞嵩草(Kobresia woodiiNoltie), 和产于西藏东南部的1 个《中国植物志》未收录的种--假钩状嵩草(K.pseuduncinoidesNoltie)。     

西藏十字花科新记录植物

报道了西藏十字花科一新记录属芝麻菜属(Eruca)和一新记录种山西异蕊芥(Dimorphostemon shanxiensis R.L.Guo et T.Y.Cheo).     

西藏睡菜科一新记录属——莕菜属

报道西藏一个新记录属,是睡菜科(Menyanthaceae)的莕菜属(Nymphoides),并报道该地区的一个新记录种,即莕菜(Nym phoides peltatum(Gmel.)O.Kuntze).     

西藏被子植物新记录

报道了西藏被子植物3种新记录,即石竹科(Caryophyllaceae)的坚硬女娄菜(Silene firma Sieb.et Zucc.)、川续断科(Dipsacaceae)的绿花刺参(Morina chlorantha Diels)和菊科(Asteraceae)的粘毛香青[Anaphalis bulley-ana(J.F.Jeffr.)Chang].     

国产悬钩子属绢毛亚组新资料

报道了采自西藏墨脱县悬钩子属绢毛亚组Rubus subsect.Lineati (Focke) Yü et Lu两个中国新纪录种,即酒红悬钩子R.calophyllus Clarke.和炫丽悬钩子R splendidissimus Hara.     

国产悬钩子属绢毛亚组新资料

报道了采自西藏墨脱县悬钩子属绢毛亚组Rubus subsect. Lineati (Focke) Yü et Lu两个中国新纪录种, 即酒红悬钩子R.calophyllus Clarke.和炫丽悬钩子R.splendidissimus Hara.     

西藏兰科植物2新记录种

报道西藏兰科2新记录种——拟毛兰Mycaranthes floribunda(D.Don)S.C.ChenJ.J.Wood和厚叶苹兰Pinalia pachyphylla(Averyanov)S.C.ChenJ.J.Wood。两种新记录植物均发现于西藏自治区墨脱县,凭证标本保存于福建农林大学林学院植物标本馆(FAFU)。     

西藏蕨类植物增补(Ⅱ)

报道了8种蕨类植物为西藏新记录,并讨论了西藏蕨类植物的区系特点和分类问题.     

西藏植物分布新资料

首次报道了西藏2个新记录属,蔷薇科(Rosaceae)的羽衣草属(Alchemilla)和鸢尾科(Iridaceae)的庭菖蒲属(Sisyrinchium),并报道2个新记录种,即无毛羽衣草(Alchemilla glabra Neygenf.)和庭菖蒲(Sisyrinchium rosulatum Bickn.)。     

中国淡水中心纲硅藻研究

本文仅报道属于中心纲硅藻的二新种和一新变种以及对菱形小环藻Cyclotella rhomboideo-elliptica Skuja(1937)特征的补充和修改。 1.罗兹直链藻西藏变种 (新变种) Melosira roeseana var.xizangensis     

武陵山区蕨类植物研究

武陵山区蕨类植物资源丰富,现知有644种(包括变种),分属116属, 46科;蕨类区系属热带-亚热带山地类型;属的地理成份有12种类型,与日本、印度-马来西亚有密切关系;武陵山区蕨类区系属我国西南区系,与华中华东区关系密切,华南、华北区次之;蕨类垂直分布分四带。     

The formation and differentiation of the Leguminosae flora in Xizang (Tibet)

Xizang (Tibet) is rich in Leguminosae flora, comprising 41 genera and 254 species so far known, exclusive of the commonly cultivated taxa (including 11 genera and 16 species). There are 4 endemic genera (with 8 species), 10 temperate genera (with 175 species) and 19 tropical genera (with 46 species) as well as the representatives of those genera whose distribution centers are in East Asia-North America, Mediterranean and Central Asia. 1. There are altogether 4 endemic genera of Leguminosae in this region. According to their morphological characters, systematic position and geographical distribution, it would appear that Salweenia and Piptanthus are Tertiary paleo-endemics, while Straceya and Cochlianths are neo-endemics. Salweenia and Piptanthus may be some of more primitive members in the subfamily Papilionasae and their allies are largely distributed in the southern Hemisphere. The other two genera might have been derived from the northern temperate genus Hedysarum and the East Asian-North American genus Apios respectively, because of their morphological resemblance. They probably came into existanc during the uplifting of the Himalayas. 2. An analysis of temperate genera There are twelve temperate genera of Leguminosae in the region, of which the more important elements in composition of flora, is Astragalus, Oxytropis and Caragana. Astragalus is a cosmopolitan genus comprising 2000 species, with its center distribution in Central Asia. 250 species, are from China so far known, in alpine zone of Southwest and Northwest, with 70 species extending farther to the Himalayas and Xizang Plateau. Among them, there are 7 species (10%) common to Central Asia, 12 species (15.7%) to Southwest China and 40 species (60%) are endemic, it indicates that the differentiation of the species of the genus in the region is very active, especially in the subgenus Pogonophace with beards in stigma. 27 species amounting to 78.5% of the total species of the subgenus, are distributed in this region. The species in the region mainly occur in alpine zone between altitude of 3500—300 m. above sea-level. They have developed into a member of representative of arid and cold alpine regions. The endemic species of Astragalus in Xizang might be formed by specialization of the alien and native elements. It will be proved by a series of horizontal and vertical vicarism of endemic species. For example, Astragalus bomiensis and A. englerianus are horizontal and vertical vicarism species, the former being distributed in southeast part of Xizang and the latter in Yunnan; also A. arnoldii and A. chomutovii, the former being an endemic on Xizang Plateau and latter in Central Asia. The genus Oxytropis comprises 300 species which are mainly distributed in the north temperate zone. About 100 species are from China so far known, with 40 species extending to Himalayas and Xizang Plateau. The distribution, formation and differentiation of the genus in this region are resembled to Astragalus. These two genera are usually growing together, composing the main accompanying elements of alpine meadow and steppe. Caragana is an endemic genus in Eurasian temperate zone and one of constructive elements of alpine bush-wood. About 100 species are from China, with 16 species in Xizang. According to the elements of composition, 4 species are common to Inner Mongolia and Kausu, 4 species to Southwest of China, the others are endemic. This not only indicates that the species of Caragana in Xizang is closely related to those species of above mentioned regions, but the differentiation of the genus in the region is obviously effected by the uplifting of Himalayas, thus leading to the formations of endemic species reaching up to 50%. 3. An Analysis of Tropical Genera There are 19 tropical genera in the region. They concentrate in southeast of Xizang and southern flank of the Himalayas. All of them but Indigofera and Desmodium are represented by a few species, especially the endemic species. Thus, it can be seen that they are less differentiated than the temperate genera. However, the genus Desmodium which extends from tropical southeast and northeast Asia to Mexio is more active in differentiation than the other genera. According to OhaShi_,s system about the genus in 1973, the species of Desmodium distributed in Sino-Himalaya region mostly belong to the subgenus Dollinera and subgenus Podocarpium. The subgenus Dollinera concentrates in both Sino-Himalaya region and Indo-China with 14 species, of which 7 species are endemic in Sino-Himalaya. They are closely related to species of Indo-China, southern Yunnan and Assam and shows tha tthey have close connections in origin and that the former might be derived from the latter. Another subgenus extending from subtropical to temperate zone is Podocarpium. Five out of the total eight species belonging to the subgenus are distributed in Sino-Himalaya and three of them are endemic. An investigation on interspecific evolutionary relationship and geographic distribution of the subgenus shows that the primary center of differentiation of Podocarpium is in the Sino-Himalaya region. Finally, our survey shows that owing to the uplifting of the Himalayas which has brought about complicated geographic and climatic situations, the favorable conditions have been provided not only for the formation of the species but also for the genus in cer-tain degree.     

横断山区唇形科植物的地理分布

横断山区唇形科植物相当丰富,有46属240种,在国内仅次于云南和四川,但明显多于其它省区。通过对其属、种的分布区类型分析,其地理分布表现如下一些特征;1)从属的分布区类型来说,横断山区唇形科植物主要是温带性质,温带分布区类型的属数占总属数(世界分布的不计算在内,下同)85.3%,其中东亚分布类型为数最多,占总属数34.1%,其次是旧世界分布类型和北温带分布类型,分别占总属数29.2%和12.2%,而东亚分布类型中占绝对优势的是中国喜马拉雅分布亚型。2)从种的分布类型来说,我国特有分布的种占绝大多数,占总种数75%,而温带分布类型和热带分布类型分别占总种数20%和5%。在我国特有分布的种中横断山区特有的占72.8%。在横断山区特有分布的种中滇西北一地特有的占横断山区特有总种数32.1%,川西南一地特有的占横断山区特有总种数9.9%,滇西北和川西南两地共同特有的占横断山区特有总种数25.2%,要是把滇西北川西南看成一整体无疑是横断山区特有种分布中心,占横断山区特有总种数67.2%。3)滇西北川西南特有种分布中心,从其特有种组成分析,其成因有历史的也有生态的,但生态成因多于历史成因,因此该中心的植物区系较为年青,这是由于新构造运动强烈、垂直气候带变化明显,冰川多次进退,导致气候上下位移频繁以横断山脉的纵向深切,促进植物在发展过程中强烈分化的结果。     

Study on the Floristic Phytogeographical Differentiation of Xizang Tibet

The geoecological conditions of Xizang (Tibet) are very complicated. An approach on the floristic phytogeographical differentiation of Xizang has been made in the present paper with a quantitative floristic method. According to areal types of the species the flora of Xizang may be classified under five major geoelements: the north temperate zone geo-element (N), the Central Asiatic geoelemcnt (C), the Qinghai-Xizang Tibetan Plateau geo-element (T), the Sino-Himalayan geo-element (SH) and the tropical geo-element (Tr). Different diagrams of spectra of floristic elements of Xizang are presented. Four cross-sections were chosen for illustrating the regional differentiation of spectra of floristic elements of Xizang. It is obvious that the SH-geo-element prevails in the Eastern and Southeastern Xizang and the Trgeo-element is confined at lower elevation of the southern flanks of the Himalayas. On the contrary, on the Plateau proper the Tgeoelement dominates and the C-geo-element plays a significant role in the northwestern part of Xizang. It corresponds to the following horizontal zones of vegetation from southeast to northwest: montane forest-alpine meadow-alpine steppe-alpine desert. An example at the southern slopes of the Eastern Himalayas has been taken to investigate the vertical variation of the spectra of floristic elements, the boundary between the both subbelts of the montane evergreen broad-leaved forest belt at an elevation of 1,800 m has been proposed as the upper limit dominated by the tropical geo-element. On the basis of dominance spectra of the fioristic elements in the grid-square system floristic boundaries are defined, which separate different floristic regions from one another, thereafter a floristic division of Xizang has been discussed. There are the sub-region of the "Himalayan flanks belonging to the Indo-Malaysian sub-kingdom of the Palaeotropical kingdom, the Sino-Himalayan sub-kingdom and the Qinghai-Xizang Plateau sub-kingdom of the Holarctic kingdom.     

秦岭落叶阔叶林25 ha森林动态监测样地物种组成与群落特征

秦岭落叶阔叶林是温带-亚热带过渡区保存较好的植被类型, 群落结构复杂。为了研究该过渡区植被的生物多样性维持机制, 按照CTFS (Center for tropical Forest Science)和中国森林生物多样性监测网络(CForBio)的建设标准, 于2015年在陕西省佛坪国家级自然保护区的落叶阔叶林建立了一块面积25 ha的动态监测样地。本文对样地内胸径(DBH) ≥ 1 cm的所有木本植物进行了调查和统计, 分析了其物种组成、区系特征、径级结构和空间分布格局。结果表明: 样地内DBH ≥ 1 cm的木本植物独立个体有47,739株, 隶属于36科66属119种。温带区系成分的科和属分别占总科数和总属数的41.18%和60.00%, 温带区系特征明显。稀有种有51种, 占总树种的42.86%。落叶树种有106种, 占总树种的89.07%, 在样地内占绝对优势。重要值≥ 1的物种共有20个, 分别占样地总个体数和总胸高断面积的70.35%和57.41%, 重要值最大的物种分别是乔木层的锐齿栎(Quercus aliena var. acutesrrata)、亚乔木层的水榆花楸(Sorbus alnifolia)和灌木层的刺叶栎(Q. spinosa)。样地内所有个体的平均胸径为8.61 cm, 整体径级分布呈倒”J”型, 小径级个体较多, 群落更新良好。除了太白杨(Populus purdomii), 样地内优势种表现出大尺度的聚集分布, 且它们的分布与生境紧密相关, 不同物种表现出不同的生境偏好。     

The Geographical Distribution and Floristic Composition of the Orchids in the Gongga Mountain Region,Sichuan

The Gongga Mountain Region, located on the eastern fringe of QinghaiXizang Plateau and at the north-eastern end of Hengduan Mountains, is one of the well-known large mountain areas in Sichuan Province. There are forty five high peaks with the elavation of 6000 m or over in the area, among which the Gongga Mountain is the highest one, with its summit being at the altitude of 7556 m, whereas the Dadu River Valley in the eastern part of the area is only 1150 m above sea level; The relative height in the region is thus about 6400 m. As we know so far the orchids in Gongga Mountain Region comprise 34 genera and 75 species with 1 variety, of which 12 species are epiphytes, 59 species with 1 variety terrestrials and 4 species saprophytes (Table 1.). I. The geographical distribution in the Gongga Mountain Region. The vertical distribution of the orchids in the area. Eastern flank: There are 39 species with I variety of orchids at 1150-2300 (-2400) m alt. in the subtropical evergreen broadleaf forest zone, of which 12 species, such as Bulbophyllum andersonii, Dendrobium hancockii, Otochilus porrecta, etc., are epiphytic (including a semiepiphyte, Pleione bulbocodioides), 25 species with 1 variety, such as Bletilla formosana, Calanthe davidii, Cypripedium henryi, etc., are terrestrial, and 2 species are saprophytic (i.e. Gastrodia elata and Neottia listeroides), the upper limit of the real epiphytic orchids is 1800 m alt. At the altitude 2300(-2400)-3600 (-3800) m, in the coniferous forest zone, found are 23 species, including 20 terrestrial species with 1 variety, such as Amitostigma gonggashanicum, Calanthe fimbriata, Coeloglossum viride, etc., 2 saprophytic species (Neottia acuminata and Risleya atropurpurea) and one semiepiphytic species. There are only 2 terrestrial species (i.e. Cypripedium tibeticum and Orchis chusua) at 3600(-3800)-4000m alt., in alpine shrub zone, the upper limit of the terrestrial orchids being 4000 m alt. At 4000-4400 m alt., in alpine meadow zone no orchids have so far been found. Western flank: There are 14 species of orchids, such as Habenaria limprichtii, Hemipilia flabellata, Satyrium ciliatum, etc., at 2300-2800 m alt., in the river valley shrub zone, and they all belong to the terrestrial orchids. At the altitude of 28003800m, in the coniferous forest zone found are 19 species of orchids, among which 18 species are terrestrial (such as Amitostigma monanthum, Cephalanthera longifolia, Peristylus coeloceras, etc.) and one is saprophytic. There are only 4 species of terrestrial orchids (i.e. Cypripedium tibeticum, Gymnadenia orchidis, Orchis chusua and O. diantha) at 3800-4800 m alt., in the alpine shrub-meadow zone, the upper limit of vertical distribution of terrestrial orchids being 4400 m alt. Twelve species here are common to the eastern flank. II. The floristic composition of orchids in Gongga Mountain Region. 1. Twenty three species, belonging to 13 genera, are widespread in Whole EastAsian Region. 2. Eight species, belonging to 8 genera, are the elements of the Sino-Japanese Subregion. 3. Forty one species with 1 variety, belonging to 24 genera, are the elements of the Sino-Himalayan Subregion, more than five times the elements of the Sino-Japanese Subregion. 4. The floristic features of the orchids in the area. (1) The floristic elements are mainly temperate and subtropical ones. (2) The life form is mostly terrestrial. (3) The species endemic to China are prolific (35 species with 1 variety, belonging to 18 genera, are endemic to China, and 26 species with 1 variety are distributed in south-western China and its adjacent region; Amitostigma gonggashanicum is endemic to the area). The floristic composition of orchids in the area is characterized by the dominance of terrestrial species and temperate and subtropical East-Asian elements, though with a few Indo-African tropical elements (such as genus Satyrium (1 species) ). In conclusion, it may be considered that the species of orchids are abundant and floristic elements are comparatively complex in the Gongga Mountain Region.     

湖南大围山杜鹃灌丛的群落组成及结构特征

灌丛是陆地生态系统中最重要的自然植被类型之一, 在植被演替、生物多样性保护和维持生态平衡等方面具有重要作用。本文基于2012年的样地资料, 对湖南大围山杜鹃(Rhododendron simsii)灌丛的物种组成和群落结构进行了分析。结果表明: 群落内共有维管束植物58种, 隶属于36科50属, 其中木本植物19种, 草本植物39种, 缺乏木质藤本; 区系以温带性质为主, 兼受热带亚热带区系的强烈影响; 群落外貌矮平, 灌木层个体集中在1-2 m高度级; 植被分类上, 该群落归属于山地中生落叶阔叶灌丛, 隶属于温性落叶阔叶灌丛群系组; 生活型以地面芽和高位芽植物为主, 反映了亚热带中山山顶温凉湿润的气候特点; 群落各层次优势种明显, 具有较高的稳定性和均匀性; 高度级和基径级分析表明群落内中等大小个体具有保守的生活史策略, 优势种杜鹃为远期衰退种群。综合分析结果表明, 该杜鹃灌丛尚处于群落演替的中期或前中期, 最终将演替为亚热带中山常绿落叶阔叶混交林或常绿阔叶林。     

Carbon storage and potentials of the broad-leaved forest in alpine region of the Qinghai- Xizang Plateau,China

Aims
Our objective was to explore the vegetation carbon storages and their variations in the broad-leaved forests in the alpine region of the Qinghai-Xizang Plateau that includes Qinghai Province and Xizang Autonomous Region.
Methods
Based on forest resource inventory data and field sampling, this paper studied the carbon storage, its sequestration rate, and the potentials in the broad-leaved forests in the alpine region of the Qinghai-Xizang Plateau.
Important findings
The vegetation carbon storage in the broad-leaved forest accounted for 310.70 Tg in 2011, with the highest value in the broad-leaved mixed forest and the lowest in Populus forest among the six broad-leaved forests that include Quercus, Betula, Populus, other hard broad-leaved species, other soft broad-leaved species, and the broadleaved mixed forest. The carbon density of the broad-leaved forest was 89.04 Mg·hm^-2, with the highest value in other hard broad-leaved species forest and the lowest in other soft broad-leaved species forest. The carbon storage and carbon density in different layers of the forests followed a sequence of overstory layer > understory layer > litter layer > grass layer > dead wood layer, which all increased with forest age. In addition, the carbon storage of broad-leaved forest increased from 304.26 Tg in 2001 to 310.70 Tg in 2011. The mean annual carbon sequestration and its rate were 0.64 Tg·a^-1 and 0.19 Mg·hm^-2·a^-1, respectively. The maximum and minimum of the carbon sequestration rate were respectively found in other soft broad-leaved species forest and other hard broad-leaved species forest, with the highest value in the mature forest and the lowest in the young forest. Moreover, the carbon sequestration potential in the tree layer of broad-leaved forest reached 19.09 Mg·hm^-2 in 2011, with the highest value found in Quercus forest and the lowest in Betula forest. The carbon storage increased gradually during three inventory periods, indicating that the broad-leaved forest was well protected to maintain a healthy growth by the forest protection project of Qinghai Province and Xizang Autonomous Region.     

青藏高原高寒区阔叶林植被固碳现状、速率和潜力

为明晰青藏高原高寒区阔叶林植被碳储量现状及其动态变化特征, 利用森林资源清查数据和标准样地实测数据, 估算了青藏高原高寒区(青海和西藏两省区)阔叶林植被的碳储量、固碳速率和固碳潜力。结果表明: 2011年青藏高原高寒区阔叶林植被碳储量为310.70 Tg, 碳密度为89.04 Mg·hm^-2。六类阔叶林型(栎(Quercus)林、桦木(Betula)林、杨树(Populus)林、其他硬阔林、其他软阔林和阔叶混交林)中, 阔叶混交林的碳储量最大, 杨树林碳储量最小; 其他硬阔林碳密度最大, 其他软阔林碳密度最小。空间分配上碳储量和碳密度表现为: 乔木层>灌木层>凋落物层>草本层>枯死木层。不同龄级碳储量和碳密度总体表现为随林龄增加逐渐增大的趋势。阔叶林碳储量从2001年的304.26 Tg增加到2011年的310.70 Tg, 平均年固碳量为0.64 Tg·a^-1, 固碳速率为0.19 Mg·hm^-2·a^-1。不同林型固碳速率表现为其他软阔林最大, 其他硬阔林最小; 不同龄级表现为成熟林最大, 幼龄林最小。阔叶林乔木层固碳潜力为19.09 Mg·hm^-2, 且不同林型固碳潜力表现为栎林最大, 桦树林最小。三次调查期间阔叶林碳储量逐渐增加, 主要原因是近年来森林保护工程的开展使阔叶林生长健康良好。     

幕阜山脉石松类和蕨类植物多样性及生物地理学特征

通过资料收集及野外调查, 得出幕阜山脉地区共有石松类和蕨类植物26科72属261种, 具有一定的东西过渡性, 为鳞毛蕨-铁角蕨植物区系, 最大属为鳞毛蕨属(Dryopteris) (29种)。其中, 庐山的物种丰富度较高(224种), 以铁角蕨属(Asplenium)为主; 幕阜山的物种密度较大(2.09种/km ²), 以卷柏属(Selaginella)为主; 九宫山以瓦韦属(Lepisorus)为主; 三者共通种有95种, 新特有现象较丰富。该区属种分化限制明显, 表现在单种科、属及寡种科、属占总科数的60%及总属数的80%。从区系成分看, 该区科、属以热带成分为主, 科和属的热带性成分与温带性成分比值(R/T值)分别为2.6和2.3。与同纬度带山地石松类和蕨类植物属的R/T值比较, 中亚热带与北亚热带交界带的蕨类植物属的R/T值在2.18-2.36之间; 种的R/T值为0.2, 为热带成分的5倍, 表现出明显的温带性质, 是罗霄山脉植物区系温带成分的重要组成部分。该区石松类和蕨类植物区系与华东、华南植物区系联系比较紧密, 表现出华东与华南两区系成分的交汇。