西藏拉萨河流域中下游不同植被类型表土花粉组合特征与年均降水量关系的研究

为定量恢复西藏拉萨河流域全新世的古气候环境,作者对拉萨河流域中下游表土孢粉组合特征与植被以及降雨量分布的相关性进行了研究。表土花粉研究结果显示,拉萨河流域中下游表土花粉组合能较好反映样方植被群落特征。根据研究区域孢粉组合特征,拉萨河流域中下游主要划分为8个孢粉组合带,即高山沼泽草甸带、高山灌丛草甸带、亚高山灌丛带、桦木林带、河谷沙棘林带、亚高山草甸带、亚高山半荒漠灌丛带和亚高山荒漠带。花粉类型与年均降水量的相关性分析结果表明,乔灌木花粉含量、Cy/G+A比值、莎草科、蓼属均与年均降水量呈显著正相关,而草本花粉含量、禾本科、蒿属、藜科与年均降水量呈显著负相关。因此,年均降水量是影响拉萨河中下游植被类型和表土花粉组合特征的重要因素。     

新疆中天山山间盆地表土花粉谱特征

基于中天山山间盆地58个表土样品花粉谱特征,结合植物群落样方物种调查结果,分析了该区域表土花粉与现代植被分布的对应关系。结果表明:(1)中天山山间盆地表土花粉组合与现代植被分布相一致,聚类分析结果将研究区分为5个植被带,按海拔由低到高的顺序分别对应荒漠草原带、山地草原带、山地森林带、亚高山草甸带、高山流石坡植被带。主成分分析第1轴揭示,湿度是植被带形成的重要影响因素。植被带之间花粉组合差异显著,荒漠草原带藜科-蒿属花粉组合优势明显,山地草原带则表现为蒿属-禾本科-藜科花粉组合,云杉属-蒿属-禾本科-藜科花粉组合及云杉属含量25%共同指示山地森林带的存在,亚高山草甸带花粉组合表现为莎草科-斗篷草属-蒿属-禾本科-藜科,高山流石坡植被带以蒿属-藜科-莎草科-斗篷草属-禾本科花粉组合为主。(2)蒿属、藜科、禾本科、云杉属、莎草科、斗篷草属是中天山山间盆地的代表物种,可作为古环境重建的重要参考。蒿属、藜科表现出超代表性,禾本科则相反。(3)中天山山间盆地植被带表土花粉分布及含量影响因素复杂,云杉属花粉受西风环流影响较大,蒿属、藜科花粉受人类活动影响显著。     

西天山南坡表土花粉分析

通过对西天山南坡不同植被带52个表土花粉样品的鉴定,研究表土花粉组合与现代植被分布的关系,分析蒿属/藜科比值(A/C)在该区域的指示意义.结果表明:山顶至山底可以划分为5个植被带,分别为高山荒漠带、高山草甸带、草甸草原带、山地荒漠带、典型荒漠带,代表性科属为蒿属、藜科、禾本科和麻黄属,该区域不同植被类型表土样品花粉组合差异明显.西天山南坡表土花粉在一定程度上受到北坡的影响,海拔越高影响越大.A/C值随海拔升高而先升高再降低,在高山草甸区域达到最高,可以作为反映山地垂直方向气候干湿变化的有效指标.由于A/C值波动较大,利用该比值进行气候环境重建时,需注意受人类活动等因素影响所产生的高值可能带来误判.     

中国西天山北坡表土花粉与区域植被关系*

天山是横亘中国西北的著名山系之一,将新疆分为南、北两个独立的地理区域,其自身也构成了一个完整的自然地理单元.因其山体海拔高,径向跨度大,纬向延伸长,气候、土壤和植被等自然地理要素各方面的区域差异,这对花粉搬运、传播、保存与沉积产生了显著影响.本文根据对西天山北坡采集的50个表土样品花粉分析,结合野外考察资料,揭示了该区域垂直方向上表土花粉组合与现代植被之间的关系,并进一步对天山表土花粉组合进行了区域对比.研究表明:(1)西天山北坡表土花粉谱可分为5个孢粉组合带,从山麓到山顶分别为山地荒漠带、草原带、森林带、亚高山草甸带、垫状植被带.(2)当云杉属花粉含量介于25％-30％时可认为该样品位于云杉林内,森林植被带花粉组合表现为以云杉属(Picea)-蒿属(Artemisia)-藜科(Chenopodiaceae)-伞形科(Apiaceae)为优势,亚高山草甸带和草原带以蒿属-藜科-禾本科(Poaceae)-云杉属-麻黄属(Ephedra)组合为主,山地荒漠带和垫状植被带以藜科-蒿属高含量为特征.云杉属花粉代表性较好,含量与现代植被分布相一致,但受气流影响显著.麻黄属含量和分布同样受气流影响,其含量不能指示该植被在样点周围存在与否.蒿属、藜科呈超代表性,只能指示区域生态意义,而禾本科则由于花粉产量低表现出低代表性.(3)西天山北坡植被带最为复杂,向东则逐渐简化.     

四川螺髻山表土和化石孢粉揭示的环境生态意义

通过四川螺髻山（1984、2017年）表土和化石孢粉与植被的关系分析发现：孢粉组合中木本植物含量（91.0%）占绝对优势,松属、冷杉属、青冈属、常绿栎类、落叶栎类、桤木属、杜鹃花科、禾本科和蒿属为主要花粉类型;人工次生林花粉组合能很好地反映母体植被的群落特征,并可指示优势种的存在和人类活动痕迹;针阔混交林花粉组合能较好地反映群落整体特征,花粉类型能与母体植被中优势种较好地对应;常绿阔叶林、针叶林和灌丛草甸的花粉组合难以反映母体植被的群落特征。DCA表明,花粉谱的百分含量,能较好地区分人工扰动植被和天然植被,但人工次生林、常绿阔叶林和针阔混交林之间以及针叶林和灌丛草甸之间未能区分。30年前后表土孢粉组合变化明显,1984年样品以松属、青冈属和桤木属等花粉为主,而2017年采样分析中松属、桤木属花粉占绝对优势。随着人类活动的加强,部分地区松树和桤木大面积飞播或种植,表土孢粉组合对当地植被指示意义与扰动强度呈负相关关系。现生植被与表土孢粉组合的差异能为恢复古植被和古环境提供参考,但原生植被已经被破坏地区的孢粉图谱,就很难作为重建历史时期植被的根据。本研究可以为亚热带山地利用孢粉学恢复第四纪时期植被与气候,探讨人类活动与环境的关系提供理论依据和实践参考。     

太白山表土花粉和气孔器与植物类型的关系研究

通过陕西太白山13个样点表土花粉组合特征和气孔器及其与植物类型之间关系的分析,结果发现:针阔混交林花粉组合能很好地反映植物类型特征,落叶阔叶林和针叶林花粉组合能较好地与植物类型相对应,高山灌丛草甸花粉组合未能反映植物类型数量特征;主要花粉类型松属、铁杉属和桦属花粉具超代表性,胡桃属和榆属花粉具适宜代表性,落叶松属、冷杉属、杜鹃花科和槭属花粉具低代表性;DCA(Detrended Correspondence Analysis)分析表明,通过花粉数据能够较好区分不同植被类型,结合气孔器特征能够准确反映植被特征。     

江西南昌西山地区表土孢粉与植被关系研究

选取江西南昌西山地区的表土和苔藓样品做孢粉分析,得出其与植被的关系,并且初步推断西山洗药湖沼泽表土花粉的来源,为钻孔花粉来源分析提供帮助。研究结果表明:南昌西山地区表土花粉组合能够较好的反映出当地植被状况。由于沼泽对区域花粉的汇聚作用,其表土花粉谱更能代表整个区域的植被状况。沼泽表层样品中主要花粉类型松属(Pinus)、蒿属(Artemisia)花粉来源于西山地区小区域和东南部季风区大区域;杉木属(Cunninghamia)、大戟科(Euphorbiaceae)等主要来源于洗药湖沼泽周围山地;莎草科(Cyperaceae)等花粉源则为沼泽内部的植物;禾本科(Poaceae)花粉来源相对复杂。聚类分析较难区分马尾松林带和杉木林带;毛竹林带、农田带、沼泽表层带和周围山地带则可以清晰地划分出来。由主成分分析可知,影响西山地区表土花粉组合的第一因子是湿度,第二因子是人类活动。     

新疆石河子南山地区表土花粉研究

天山作为亚洲大陆最大的山系之一,横贯于新疆的中部,成为分隔南、北疆自然地理区系的山系,它对花粉的传播、保存、搬运与沉积具有重大作用。根据对西北干旱区域新疆石河子南山地区一条沿着海拔高度从2400 m到300 m的样带所采集的23个表土花粉样品的孢粉组合图式和现代植被样方调查资料,探讨了北坡垂直带的植被与表土花粉之间的关系。该区表土孢粉谱可分为4个孢粉组合带,分别对应森林植被带、森林草原植被带、蒿属荒漠带和典型荒漠带。比较特殊的是典型荒漠带被划分为两个亚带,一个亚带是以蒿属、藜科占主要成分的典型荒漠带,另一亚带蒿属、藜科含量较高并含有大量沼泽蕨和芦苇植硅体,兼具典型荒漠和湿地特征。在海拔400 m以上,孢粉组合与现代植被的对应关系较好,带Ⅰ中较高含量的云杉花粉验证了以云杉为主的森林植被带。带Ⅱ中以云杉为主的乔木植物和含量较高的藜科、蒿属和蓼属等草本植物为主的孢粉组合特征与森林-草原植被带的植被特征较为类似。云杉属花粉在海拔低于1350 m的地方即林带下方所占的比例很小,一方面由于距林地的距离较远,另一方面,可能是山风气流对云杉花粉往下搬运的能力较弱所致。带Ⅲ的蒿属花粉含量较高,与该带植被中绢蒿较多有一定的关系,带Ⅳ以藜科为主的花粉组合特征代表了这个植被带的荒漠植被类型。但是在海拔400 m之下,带Ⅳ的亚带Ⅳ₂的高含量的沼泽蕨和芦苇植硅体的孢粉组合在一定程度上还代表了古湿地环境。通过该部分表土花粉组合特征与草滩湖剖面孢粉谱的对比,验证了当地农业种植选址的生态可行性,同时就开垦程度对环境的影响进行了初步探讨。另外,亚带Ⅳ₂的蒿属/藜科(Artemisia/Chenopodiaceae(A/C))比值比亚带Ⅳ₁高,可能与该样点受人为扰动较大有关。     

新疆赛里木湖流域表土花粉组合与现代植被关系研究

通过对新疆赛里木湖流域现代植被调查及不同植被带的53个表土样品花粉鉴定分析,深入研究了该区域表土花粉组合与现代植被的关系。研究结果表明:表土花粉组合可反映出该流域植被的总体特征,赛里木湖流域不同植被带内花粉组合差异显著,依据花粉谱特征可以分为草原带、针叶林带、森林灌丛带、亚高山草甸带、荒漠草原带、草原灌丛带、荒漠植被带;云杉属、蒿属、藜科、禾本科含量高、变化明显,可作为该流域古环境重建的重要参考依据;蒿属/藜科(A/C)比值可以用来区分荒漠植被带和荒漠草原带,荒漠植被带A/C值平均为0.62,荒漠草原带A/C值平均为1.55,1是两者之间的界值;云杉属花粉含量≥30%时可认为该样品来自于云杉林,云杉属-蒿属-禾本科等花粉组合表明森林(灌丛)的存在,亚高山草甸以蒿属-禾本科-云杉属组合为主,(荒漠)草原(灌丛)以蒿属-藜科-禾本科为主,藜科-蒿属是荒漠植被的建群种。     

东北长白山垂直林带下现代表土花粉与植被关系

在长白山北坡5个垂直植被带的林下采集29个表土(苔藓)样品进行孢粉分析,同时将其结果作对应分析.结果表明,除个别样品,在长白山垂直植被带的孢粉组合中乔木花粉占优势,其中以松属和桦木属最多,在各植被带均有分布.松属花粉最高含量出现在以红松为主的针阔混交林带的孢粉组合中;桦木属花粉则大量见于亚高山岳桦林带和以栎桦为主的阔叶落叶林带,但前者伴有耐寒、早的小灌丛杜鹃花属花粉,而在后者中则见有大量蕨类孢子.阔叶落叶林带的孢粉组合是以类型多、百分含量高的落叶乔木花粉和大量蕨类孢子为特征.亚高山针叶林带的孢粉组合中出现数量众多的云杉属花粉.在高山苔原带孢粉组合中虽然乔木花粉略占优势,但极少或不见蕨类孢子,与其他植被带比较,草本植物和小灌丛花粉明鲜较多,其中杜鹃属花粉含量在草本中占优势,其中出现苔原带的指示花粉--仙女木属.     

Numerical Characteristics of Pollen Assemblages of Surface Samples from the West Kunlun Mountains

A total of 31 suface sediment samples were collected from West Kunlun Mountain in south Xinjiang Autonomous Region in northwest China. These samples are from seven types of vegetation: Picea schrenkiana Fisch. et Mey. forest, Sabina Spach. woodland, sub-alpine steppe, alpine meadow, desert vegetion, cushion-vegetation and vegetation adjancent to glaciers. Pollen percentages and pollen concentrations were calculated in all samples. The dominant pollen types in the region are Chenopodiaceae, Artemisia, Picea, Ephedra, Gramineae, Cyperaceae, Rosaceae, Leguminosae, Compositae etc. In order to reveal the relationship between pollen composition and the vegetation type from which the soil sample was collected, principal component analysis and group average cluster analysis were employed on the pollen data. The results revealed that the major vegetation types in this region could be distinguished by pollen composition: a. Samples from desert vegetation were dominated by pollen of Chenopodiaceae (about 60195%). The percentages of all other pollen types were low. b. Picea forest samples were rich in Picea pollen (about 20%) Sabina forest had more Sabina pollen grains than other vegetation types (about 5%, others <1%). Pollen percentages of Artemisia, Chenopodiaceae and Ephedra were comparatively higher (each about 20%) in these samples from the two types of vegetations. C. Pollen percentages of Artemisia, Cyperaceae, Gramineae and Chenopodiaceae were high in both sub-alpine steppe and alpine meadow. But steppe containal more Artemisia and Chenopodiaceae (steppe 33.75% and 32.30%, meadow 15.57% and 19.48% in average), less Cyperaceae and Gramineae (steppe 2.58% and 7.60%, meadow 22.35% and 12.93% in average) than meadow. d. Samples from cushion-vegetation and vegetation adjacent to glaciers were mainly composed of pollen grains transported from other sites. It was not easy to distinguish them from other vegetation types. Principal component analysis and cluster analysis distinguish samples from Picea forest, Sabina woodland, sub-alpine steppe, alpine meadow and desert vegetation. Therefore we think it will be possible to apply the module to reconstruct past vegetation in this region and other similar regions. Regression analysis was also applied to reveal the relationships between pollen and plant percentages of Artemisia, Chenopodiaceae, Cyperaceae and Gramineae. The results indicated that a linear relationship existed between pollen and plant percentages for Artemisia, Chenopodiaceae and Cyperaeeae.     

干旱区石羊河流域河水孢粉组合特征

石羊河流域初春和仲夏两季不同地点 ,洪水期、平水期 30个河水样孢粉分析显示 ,没有人类影响的情况下 ,河水样孢粉组合是由河流上游径流区的植物孢粉和采样点附近植被孢粉组成的 ,前者在孢粉组合中至少占30 %～ 6 0 %。河水搬运孢粉的能力非常强 ,可长距离、大量地搬运孢粉。孢粉组合中河水贡献孢粉的含量较高 ,如 :云杉属 (PiceaDietr.)花粉河水的贡献率在中下游可达 16 .5 %和 7.7%。采样时间、地点影响河水的孢粉组合和浓度 ,洪水期是孢粉搬运的主要时期。河水经过水库后 ,其中孢粉绝大多数沉积在水库中。     

燕山地区表土花粉与植被间的数量关系

 根据表土花粉分析和植被样方调查,燕山地区各植被带的表土孢粉组合在总体面貌上代表了植被组成,但孢粉组合不等于植被组成,而影响其差异的原因是孢粉保存、孢粉产量、外来孢粉和孢粉鉴定。表土花粉的代表性因种属不同而差异颇大,乔木植物花粉中松(Pinus)、桦(Betula)、胡桃     

燕山南部花粉散布特征

燕山南部花９粉用布特性研究表明,空气中的花粉反映的是植物花期的季节性变化,表 粉是植物多年花粉散布的混合,冲积物花粉与空气中和表土花粉显著不同,是取样点上游表土花粉的混合,与取样点周围植被关系较小。因此,根据冲积物花粉恢复古植被,反映的是整个流域的植被面貌,而不是某一点的植被特征。     

阴山山脉东段花粉通量及其与表土花粉比较研究

阴山山脉东段蛮汉山和大青山山顶草甸、桦木林、虎榛子灌丛、人工油松林及杂草草原5个植被类型花粉通量与表土花粉对比研究发现,不同植被类型及同一植被类型不同样点花粉通量和花粉浓度差异明显,捕捉器样品花粉源区面积小于表土样品,能较好地反映样品点周围植被组成,但受虫媒植物和局地植物花粉影响,与表土样品花粉组合差异明显;表土样品中松属、桦属、蒿属及藜科花粉含量多高于捕捉器样品,表明这些花粉不仅易于传播,而且在表土中具有较强的保存能力;禾本科花粉具低代表性,花粉保存能力较低.地层花粉分析中应引起注意.     

Preliminary study on pollen distribution in the surface soil of the Turpan region in the southern slope of Tianshan Mountains,Xinjiang, China

Aims Our main purposes were to analyze the relationship between vegetation and pollen in the surface soil of the Turpan region, which is located in the southern slope of the eastern Tianshan Mountains, and to compare different pollen assemblages between the Turpan region and the northern slope of the Tianshan Mountains.Methods We collected 36 modern pollen samples and carried out modern vegetation survey in the Turpan region along an altitudinal gradient from 2 000 m to -154 m. Detrended correspondence analysis and Redundancy analysis were applied to analyze the distribution pattern of pollen in surface soils. Important findings We divided the pollen spectra into four pollen assemblage zones (mountain desert-steppe and desert, Gobi gravel, typical desert and salt mash vegetation), corresponding to the major vegetation types in the Turpan region. When compared with the northern slope, the characteristics of pollen assemblages in the mountain desert-steppe and desert were similar to those in the forest-steppe on the northern slope of the Tianshan Mountains; the pollen assemblages in the Gobi gravel and the typical desert seemed to be more consistent with those in the typical desert on the northern slope; however, no analogue was found in the salt mash vegetation. Obviously, the vertical pollen spectra in Turpan region were incomplete, lacking typical forest and Artemisia desert pollen zones. Besides, similar pollen zones in the Turpan region were found at an elevation of about 300 m higher than those in the northern slope. It is remarkable that the typical tree pollen, such as Picea and Pinus, showed their extra representation in the Turpan region. On one hand, the valley forest on the southern slope of the Tianshan mountains played an important role in pollen dispersal. On the other hand, with the cold air on the northern slope over the Tianshan Mountains, pollen may be carried and deposited in the Turpan region. The rivers feeding into Aiding Lake in the Turpan region may also contribute to the distribution of Picea and Pinuspollen. Lots of pollen studies have shown that the ratio of Artemisia to Chenopodiaceae (A/C) can be used as a good indicator of the degree of humidity in the semi-arid and arid areas. Our study revealed that A/C can roughly reflect the characteristics of the desert zone in the study area. The results of Redundancy Analysis ordination on pollen assemblages and environmental factors (mean annual temperature (MAT), mean annual precipitation (MAP) and altitude (ALT)) revealed that MAP was the main environmental factor affecting the pollen assemblages in the surface soil in the Turpan region and had more significant effects on the distribution of Nitraria pollen than on the distribution of Artemisia and Chenopodiaceae.     

甘肃兴隆山国家自然保护区表土花粉组合及数量分析

提要通过对兴隆山国家自然保护区不同植物群落40个样点的表土花粉组合特征、数量分析[分层聚类分析（HCA）和降趋势对应分析（DCA）],研究了花粉组合与植物群落的关系。结果显示：1）大部分森林和部分灌丛的花粉组合与植物群落之间对应关系很好,如青杆林（云杉属组合）、辽东栎林（栎属-桦属-蒿属组合）、桦林（桦属-蒿属-沙棘属组合）和沙棘灌丛（沙棘属组合）;2）蔷薇灌丛（蒿属-蔷薇科组合）和草原（蒿属-菊科组合）的花粉组合基本上能反映群落特征;3）山杨林、糙皮桦灌丛和高山灌丛草甸的花粉组合与植物群落的对应性较差;4）云杉属花粉的长距离传播能力有限;5）栎属花粉仅在辽东栎林中含量较高;6）桦属花粉含量较高时（〉30％）能够反映桦林的存在,含量较低时（〈10％）可能当地无桦树生长;7）沙棘属和蒿属花粉具超代表性;8）豆科、禾本科和蔷薇科花粉具低代表性;9）HCA和DCA分析对于区分主要花粉类型和揭示环境梯度是有效的,但反映植被细节特征的能力有限。     

Monitoring the fluvial palynomorph load in a lowland temperate catchment and its relationship to suspended sediment and discharge

Despite it being a component of the seston we know very little about fluvial (waterborne) pollen and spore (palynomorph) transport. This paper presents the results of a monitoring programme conducted over two years and at a catchment scale in South West England. A hierarchical monitoring network was established with flood peak samples taken at 9 sub-catchments, intra-hydrograph samples taken in two sub-catchments and time-integrated sampling undertaken at one location. In addition sampling was undertaken of probable palynomorph sources such as channel bed and bank sediments, and the airborne pollen flux was monitored using modified Tauber traps. The results support previous research in illustrating how the vast majority of fluvial pollen and spores are transported during floods (91%) and that the main control on waterborne palynomorph assemblages is the catchment vegetation and its spatial distribution but with a long-distance (extra-catchment) component. However, strong seasonal effects are also shown, and the importance of distinctive sources such as the riparian input, bed re-suspension and overland flow into drains and tributaries is revealed. Fine sediment in river pools appears to act as a selective store of damaged cereal type pollen grains derived from arable fields. Although pollen does form part of composite particles the data presented here suggest that the majority of the pollen is transported as single grains. Fluvial palynomorph loading is strongly dependant upon discharge and so concentrations in laminated or varved sediments could be regarded as a proxy for flood magnitude. Handling editor: J. Saros