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Nektaspid arthropods from the Lower Cambrian Emu Bay Shale Lagerstätte,South Australia,with a reassessment of lamellipedian relationships
Authors:JOHN R PATERSON  GREGORY D EDGECOMBE  DIEGO C GARCÍA‐BELLIDO  JAMES B JAGO  JAMES G GEHLING
Institution:1. Division of Earth Sciences, School of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia;2. e‐mail jpater20@une.edu.au;3. Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, U.K.;4. e‐mail g.edgecombe@nhm.ac.uk;5. Departamento de Paleontología, Instituto de Geología Económica (CSIC‐UCM), Facultad de Ciencias Geológicas, 28040‐Madrid, Spain;6. e‐mail Diego.GBC@geo.ucm.es;7. School of Natural and Built Environments, University of South Australia, Mawson Lakes, SA 5095, Australia;8. e‐mail jim.jago@unisa.edu.au;9. South Australian Museum, North Terrace, Adelaide, SA 5000, Australia;10. e‐mail gehling.jim@saugov.sa.gov.au
Abstract:Abstract: The lower Cambrian Emu Bay Shale on Kangaroo Island, South Australia, contains the only known Cambrian Burgess Shale‐type biota in Australia. Two new lamellipedian arthropods, Emucaris fava gen. et sp. nov. and Kangacaris zhangi gen. et sp. nov., from the Emu Bay Shale Lagerstätte are described as monotypic genera that are resolved cladistically as a monophyletic group that is sister to Naraoiidae + Liwiidae and classified within the Nektaspida as a new family Emucarididae. Shared derived characters of Emucarididae involve a bipartite, elongate hypostome and elongation of the pygidium relative to the cephalic shield and very short thorax. A monophyletic Liwiidae is composed of Liwia and the Ordovician Tariccoia + Soomaspis but excludes Buenaspis, and even the membership of Buenaspis in Nektaspida is contradicted amongst the shortest cladograms. New morphological interpretations favour affinities of Kwanyinaspis with Conciliterga rather than with Aglaspidida, and Phytophilaspis with Petalopleura.
Keywords:Arthropoda  Lamellipedia  Nektaspida  Emucaris  Kangacaris  phylogeny
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