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101.
Fishes are often thought to have passed through mass extinctions, including the Cretaceous-Tertiary (KT) event, relatively unscathed. We show that neoselachian sharks suffered a major extinction at the K/T boundary. Out of 41 families, 7 became extinct (17±12%). The proportional measure increases at lower taxic levels: 56±10% loss of genera (loss of 60 out of 107) and 84±5% loss of species (loss of 182 out of 216). However, the Maastrichtian and Danian are characterized by a high number of singleton taxa. Excluding singletons we have calculated a 34±11% loss of genera and a 45±9% loss of species. The simple completeness metric (SCM) for genera displays a decrease from the Maastrichtian (94%) to the Danian (85%) indicating a rather complete fossil record of neoselachian genera. The extinctions were heavy among both sharks and batoids (skates and rays), but most severe among batoids, which lost almost all identifiable species. There were equal losses among open marine apex predators (loss of Anacoracidae, Cretoxyrhinidae, and Scapanorhynchidae) and durophagous demersal forms from the continental shelf and shallow seas (Hypsobatidae, Parapaleobatidae, Sclerorhynchidae, Rhombodontidae). Benthopelagic and deep-sea forms were apparently little affected. New families with similar ecological roles (Carcharhinidae, Isuridae, Torpedinidae) replaced these families in the Danian, and full diversity of the different shark and batoid groups had been recovered by the end of the Paleocene or early Eocene. Sharks and rays suffered levels of extinction entirely in line with other groups of organisms at the K/T extinction event.  相似文献   
102.
Integrating fluxes from heterogeneous vegetation   总被引:1,自引:0,他引:1  
The vegetated landscape of Europe has been strongly impacted by human management to produce a heterogeneous patchwork of semi‐natural and agricultural vegetation varying over a wide range of spatial scales. A model is described for averaging vegetation fluxes from a landscape of forest and grassland into the planetary boundary layer (PBL). At a scale of 1 km, model simulations indicate that vegetation heterogeneity exerts little effect on the PBL and regional fluxes will be simple areal averages of the different vegetation types. Above 5 km the model simulates significant effects of different vegetation types on the whole PBL. Averaging fluxes to the regional scale will therefore need to consider explicitly the nature, extent and behaviour of different vegetation types.  相似文献   
103.
景观边界研究概况   总被引:13,自引:3,他引:10  
常禹  布仁仓  胡远满 《生态学杂志》2002,21(5):49-53,F004
景观边界指在特定时空尺度下 ,相对均质的景观之间所存在的异质景观[6 9] 。过去 ,生态学家的注意力往往集中在景观边界两侧的同质区域来研究生态系统的结构和过程 ,忽视生态系统间的异质区域 ,它们在地图上往往被忽视或缩成一条线。但是 ,景观边界是一个客观存在的空间实体 ,有自己特定的结构和功能。由于景观边界包含有不同类型和不同尺度上的均质景观和异质景观 ,因而它具有等级结构[13] ,同时景观边界都有长度和宽度 ,在此范围内 ,相邻景观单元的小斑块交叉出现 ,因此景观边界是相邻景观单元相互转化的发生区。从景观的演替过程可以看出…  相似文献   
104.
关于"一类多分子生化反应系统的极限环"一文的补充   总被引:2,自引:1,他引:1  
本文用构造无穷大外境界线的方法应用Poincare-Bendixson环域定理解决了系统x=δ-xyn;y=xyn-ay/(y+b)存在极限环的充要条件,从而与文[1]共同完整地解决了此系统的定性分析.  相似文献   
105.
The Gebel Qreiya and nearby Wadi Hamama sections of the central Eastern Desert are among the most complete K/T boundary sequences known from Egypt. The two sections were analyzed spanning an interval from l.83 Myr below to about 3 Myr above the K/T boundary. A 1-cm-thick red clay layer at the K/T boundary at Gebel Qreiya contains an Ir anomaly of 5.4 ppb. The high-resolution study and well-preserved nannoflora provide good age control and the first quantitative records of calcareous nannofossil assemblages for paleoecological interpretations across the K/T transition in Egypt. Four zones (Micula murus, Micula prinsii, NP1, and NP2) were distinguished and correlated with other nannofossil and planktonic foraminiferal zonations that are broadly applicable for the eastern Tethys region. Latest Maastrichtian assemblages are abundant and diverse, though Cretaceous species richness progressively decreased across the K/T boundary. Dominant species include Arkhangelskiella cymbiformis, Micula decussata and Watznaueria barnesae, with high abundance of dissolution-resistant M. decussata reflecting periods of high environmental stress. Thoracosphaera blooms mark the K/T boundary and are followed by an acme of the opportunistic survivor Braarudosphaera bigelowii, the first appearance of the new Tertiary species Cruciplacolithus primus, and an acme of Coccolithus cavus/pelagicus. These successive abundance peaks provide the basis for subdivision of the Early Danian Zones NP1 and NP2 into five subzones. Correlation of selected nannofossil taxa from the Egyptian sections with those from various onshore marine and deep-sea sections provides insights into their paleoenvironmental and paleoecological affinities.  相似文献   
106.
Development and differentiation of the vertebrate caudal midbrain and anterior hindbrain are dependent on the isthmic organizer signals at the midbrain/hindbrain boundary (MHB). The future MHB forms at the boundary between the Otx2 and Gbx2 expression domains. Recent studies in mice and chick suggested that the apposition of Otx2- and Gbx2-expressing cells is instrumental for the positioning and early induction of the MHB genetic cascade. We show that Otx2 and Gbx2 perform different roles in this process. We find that ectopically expressed Otx2 on its own can induce a substantial part of the MHB genetic network, namely En2, Wnt1, Pax-2, Fgf8 and Gbx2, in a concentration-dependent manner. This induction does not require protein synthesis and ends during neurulation. In contrast, Gbx2 is a negative regulator of Otx2 and the MHB genes. Based on the temporal patterns of expression of the genes involved, we propose that Otx2 might be the early inducer of the isthmic organizer genetic network while Gbx2 restricts Otx2 expression along the anterior-posterior axis and establishes an Otx2 gradient.  相似文献   
107.
黔南石炭-二叠系界线牙形刺序列的再研究   总被引:6,自引:3,他引:3  
本文详细描述了华南贵州罗甸纳水和紫云羊场石炭-二叠系的界线牙形刺序列,并自上而下可划分为:Streptognathodus barskovi,S.constrictus,S.isolatus,S.wabaunsensis.S.tenuialueus和S.firmus带,而Strep-tognathodus isolatus的首现则代表了二叠系的底界,本文讨论了世界各地石炭-二叠系界线牙形刺序列的对比。  相似文献   
108.
The contributions of conformational dynamics to substrate specificity have been examined by the application of principal component analysis to molecular dynamics trajectories of alpha-lytic protease. The wild-type alpha-lytic protease is highly specific for substrates with small hydrophobic side chains at the specificity pocket, while the Met190-->Ala binding pocket mutant has a much broader specificity, actively hydrolyzing substrates ranging from Ala to Phe. Based on a combination of multiconformation analysis of cryo-X-ray crystallographic data, solution nuclear magnetic resonance (NMR), and normal mode calculations, we had hypothesized that the large alteration in specificity of the mutant enzyme is mainly attributable to changes in the dynamic movement of the two walls of the specificity pocket. To test this hypothesis, we performed a principal component analysis using 1-nanosecond molecular dynamics simulations using either a global or local solvent boundary condition. The results of this analysis strongly support our hypothesis and verify the results previously obtained by in vacuo normal mode analysis. We found that the walls of the wild-type substrate binding pocket move in tandem with one another, causing the pocket size to remain fixed so that only small substrates are recognized. In contrast, the M190A mutant shows uncoupled movement of the binding pocket walls, allowing the pocket to sample both smaller and larger sizes, which appears to be the cause of the observed broad specificity. The results suggest that the protein dynamics of alpha-lytic protease may play a significant role in defining the patterns of substrate specificity. As shown here, concerted local movements within proteins can be efficiently analyzed through a combination of principal component analysis and molecular dynamics trajectories using a local solvent boundary condition to reduce computational time and matrix size.  相似文献   
109.
黔东早—中寒武世凯里组疑源类组合及其界线意义   总被引:13,自引:0,他引:13  
通过对台江八郎和丹寨两条剖面凯里组中疑源类化石的分析,研究,台江剖面产疑源类化石有17属35各(12未定种),其中以Leiosphaeridia,Synsphareidium,Cymatiosphaera,Pterospermella最为丰富,可划分为3个组合,即Cymatiosphaera cf.cristata-Fimbriaglomerella memebrancea组合,Cristallinium-Micrhystridium-Pterospermella组合和Dictyotidium-Granomarginata组合,丹寨平寨剖面凯里组所产疑源类化石有:13属21种(5未定种),同样划分为3个组合,即Leiosphaeridia-Tasmanites组合,Retisphaeridium-Micrhystridium tentatium组合和Baltisphaeridium-Bubomorpha hunjiangensis组合,本文对两条剖面中的凝源类化石的组成,相对含量做了详细的统计,发现在八郎剖面9-2层和平寨剖面3层疑源类的丰度,分异度开始发生明显的变化,表现由早寒武世向中寒武世疑源类组合面貌的转变,这一疑源类转变层位正好是与三叶虫划分的中,下寒武统界线的层位位置相一致,这充分表明疑源类化石可作为划分中,下寒武统的极有价值的微体生物化石证据。  相似文献   
110.
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