南岭国家级自然保护区兰科植物物种多样性的海拔梯度格局

兰科是单子叶植物最大的科和被子植物中的第二大科.兰科植物物种多样性的海拔梯度格局因地域或坡向不同而呈不同趋势.南岭国家级自然保护区地处南岭山脉中段南坡,植物资源丰富.作者对南岭国家级自然保护区兰科植物进行了为期4年的调查,记录了其种类、数量、生活型、海拔、生境,分析了其区系成分,并依据IUCN(2001)濒危物种红色名录及保护区的具体情况对其濒危等级进行了评估.从海拔230-1,902m,每200 m分段,共划分为9个海拔段分析其垂直分布格局.结果显示:(1)实地记录兰科植物38属70种,大多数种类数量较少,乳阳管理处的种类最为丰富;(2)随着海拔的上升,总体兰科植物以及不同生活型的兰科植物物种数量均呈现"中间高度膨胀型"而峰值偏向低海拔的特点;(3)濒危物种主要集中在600-1,000 m人为活动稀少、环境好的小生境中;(4)800-1,000 m的区系成分最丰富,热带性质的兰花多数分布在海拔1,600 m以下;(5)相邻海拔段的兰科植物种类的相似性系数较高;海拔段相隔越远,相似性系数越小.小生境和水分条件对于兰科植物的生长非常重要,中海拔具有良好的水热条件,因此物种多样性最高;而低海拔地区的人为破坏使得该海拔段兰科植物物种的多样性显著降低.     

金佛山自然保护区兰科植物多样性及保护对策研究

通过对保护区内兰科植物的调查,分析了保护区内兰科植物的物种多样性、垂直分布格局.结果表明,金佛山兰科植物种类丰富,区系成分复杂多样,共有兰科植物48属114种;金佛山兰科植物特有现象明显,中国特有兰科植物47种,3种为金佛山特有;垂直分布格局明显;不同海拔段的兰科植物濒危程度差异较大,低海拔地区物种?受威胁程度较低,资源保存状况更好.对造成兰科植物濒危的主要原因进行了分析,并提出了保护对策.     

高黎贡山种子植物物种丰富度沿海拔梯度的变化

物种丰富度沿海拔梯度的分布格局成为生物多样性研究的热点。为探讨中尺度区域物种丰富度沿海拔梯度的分布,本文以高黎贡山为研究对象,利用该地区的地方植物志资料,结合通过GIS生成的区域数字高程模型(DEM)数据,分析了该区域全部种子植物和乔木、灌木、草本三种生活型种子植物物种丰富度的垂直分布格局以及物种密度沿海拔梯度的变化特征。结果表明：(1)全部种子植物和不同生活型植物物种丰富度随着海拔的升高呈现先增加后减小的趋势,最大值出现在海拔1500—2000m的范围;(2)物种密度与海拔也呈现单峰曲线关系;(3)物种丰富度和物种密度分布格局的形成主要受海拔所反映的水、热状况组合以及物种分布的边界影响。     

井冈山自然保护区野生兰科植物资源分布及特点

通过对江西省井冈山自然保护区兰科(Orchidaceae)植物的调查,分析了保护区内兰科植物的物种多样性、区系特点、分布格局,并采用SPSS18.0软件对井冈山及其邻近保护区属的种类进行相关和聚类分析。结果表明:井冈山兰科植物种类丰富,共有兰科植物34属75种,中国特有兰科植物19种。井冈山兰科植物资源垂直分布格局明显,不同海拔段的兰科植物分布差异较大,中低海拔的物种分布较多,资源相对保存较好;区系成分复杂多样,井冈山兰科植物属的地理成分可以划分为10个类型2个变型,热带性属占优势,占总属数的67.3%,但热带分布的种仅占本地区总种数的37.3%,而温带分布的种却占62.7%,故井冈山自然保护区属热带植物区系与温带植物区系过渡的交汇地带。井冈山兰科植物与齐云山自然保护区的相似性最高,与武夷山自然保护区的相似性最低。     

鄂西北珍稀濒危植物及物种多样性空间尺度分析

为了探究鄂西北地区珍稀濒危植物及其多样性空间分布与该区域地理环境的关系,于2007-2017年先后对鄂西北8个自然保护区的植物资源进行了调查,对其种类组成、地理成分、生活型及物种多样性的空间分布特征进行研究。结果显示:(1)鄂西北珍稀濒危植物共有262种,隶属于62科149属;(2)该区域珍稀濒危植物地理成分与生活型谱具有地域过渡性和多样性;(3)鄂西北地区珍稀濒危植物的物种丰富度峰值出现在海拔900～1700 m区段,物种丰富度与海拔区段间具有显著的二次方程关系:y=-0.5107x~2-12.43x+7.3111(R~2=0.7867,P=0.04);(4)不同空间尺度下,物种Shannon-Wiener多样性指数(H_D)、均匀度指数(ED)、Margalef多样性指数(K_D)随空间尺度的变化均呈幂函数关系,其相关系数分别为R~2=0.7303 (H_D)、R~2=0.8858(E_D)、R2=0.9805(KD)。研究表明H_D、E_D、K_D值的变化幅度可能与该区域地理环境的变化有关。目前砍伐毁林和过度采挖是该地区珍稀濒危植物最大的威胁因素,建议从珍稀濒危植物物种多样性分布与地理环境特性相结合的角度加强管理,为区域珍稀濒危植物的科学恢复提供支撑。     

江西三清山兰科植物区系分析

江西省三清山兰科植物资源丰富,约21属30种,占江西省兰科植物属种的56.76%、38.46%,占中国兰科植物属种的12.28%、2.41%。区内兰科植物生活型以地生兰为主;分布区式样以温带型为主;区系成分分散,缺乏本地特有成分。与各山地的相似性以及属种分布型比较显示,三清山与5山地在属级水平上相似性系数高,兰科植物区系联系比较紧密,从属分布区类型看,三清山、黄山、天目山和庐山以北温带分布为主,井冈山和武夷山以热带亚洲-热带大洋洲为主;在种级水平上相似性系数较低,种分布区类型均以东亚和中国特有分布为主。按生态型划分,地生兰为主,附生兰为辅,三清山和黄山只有两生态类型,武夷山、天目山、井冈山和庐山生态类型齐全。最后,依据"中国物种红色名录"对各山地珍稀濒危保护物种进行归纳统计。     

白水江国家级自然保护区野生资源植物的垂直分异研究

基于先期建成的种子植物数据库,分析了白水江国家级自然保护区野生资源植物的物种丰富度、资源结构及性状特点沿海拔梯度的分异规律.结果表明:(1)根据资源植物物种数量和种数增减速率,全区可划分为4段:速增较丰富带、稳定丰富带、速减较丰富带和速减贫乏带;(2)根据各100 m海拔段之间植物资源结构的相似性进行聚类分析,结果显示药用植物种数的变化是形成一级分聚的主导因子;(3)随着海拔梯度的上升,资源植物的区系成分、性状特征表现出相对应的垂直分异规律.     

甘肃白水江国家级自然保护区珍稀植物的垂直分布及其保护

白水江自然保护区有各类珍稀濒危植物104种,占该区总种数的5.28%。其中划归《国家重点保护野生植物名录(第一批)》的有20种,划归《中国珍稀濒危保护植物名录》的有37种,划归《国家珍贵树种名录》的有18种。该区在整个海拔梯度上均有珍稀植物分布,但很不均匀。根据珍稀植物物种丰富度的高低,大体可分为“极丰富段”“较丰富段”“丰富段”“稀少段”4段。不同类型的珍稀植物的垂直分布也有差异。根据白水江珍稀濒危植物的垂直分布格局和保护现状提出了几点建议。     

广西雅长自然保护区兰科植物多样性研究

对广西雅长兰科植物自然保护区的兰科植物多样性进行调查研究。该保护区兰科植物多样性具有如下特点:(1)物种多样性丰富,居群数量大,共有兰科植物44属113种(含5变种);(2)分布广泛,生境复杂多样;(3)地理成分兼备热带和亚热带性质,生活类型齐全;(4)垂直分布格局呈现一定规律性。文章最后提出了该保护区兰科植物多样性保护及利用的初步建议。     

秦岭兰科植物地理区系特征

秦岭是中国野生兰科植物最为丰富和区系分化率最高的地区之一,但是有关该地区兰科植物的系统整理尚未见到,本研究为秦岭兰科植物资源保护提供基础数据支持,为今后秦岭国家公园建设和生物的重点区域进行了补充调查和标本采集,形成了秦岭地区兰科植物名录,依据中国种子植物分布区类型的划分方法以及GIS软件,分析它们的属种特征、区系成分以及空间分布格局。秦岭兰科植物有52属149种,其中中国特有种70种,占总种数的47%;生活型有地生型、附生型、半附生型和腐生型,以地生型为主;秦岭兰科植物主要分布在海拔800~3 400 m;热点地区为陇南西南山地、太白-佛坪山地、伏牛山地。同时,在区系分析的基础上,提出秦岭兰科植物的区系特征,发现：（1）种类丰富,温带性质显著;（2）起源古老,新老兼并;（3）特有成分繁多;（4）珍稀濒危保护植物聚集;（5）丰富度随生境的变化而不同。秦岭兰科植物地理区系研究为植物生物多样性的保护提供基础数据支持,在秦岭植物多样性研究方面具有一定的意义。     

Orchid Species Richness along Elevational and Environmental Gradients in Yunnan,China

The family Orchidaceae is not only one of the most diverse families of flowering plants, but also one of the most endangered plant taxa. Therefore, understanding how its species richness varies along geographical and environmental gradients is essential for conservation efforts. However, such knowledge is rarely available, especially on a large scale. We used a database extracted from herbarium records to investigate the relationships between orchid species richness and elevation, and to examine how elevational diversity in Yunnan Province, China, might be explained by mid-domain effect (MDE), species–area relationship (SAR), water–energy dynamics (WED), Rapoport’s Rule, and climatic variables. This particular location was selected because it is one of the primary centers of distribution for orchids. We recorded 691 species that span 127 genera and account for 88.59% of all confirmed orchid species in Yunnan. Species richness, estimated at 200-m intervals along a slope, was closely correlated with elevation, peaking at 1395 to 1723 m. The elevational pattern of orchid richness was considerably shaped by MDE, SAR, WED, and climate. Among those four predictors, climate was the strongest while MDE was the weakest for predicting the elevational pattern of orchid richness. Species richness showed parabolic responses to mean annual temperature (MAT) and mean annual precipitation (MAP), with maximum richness values recorded at 13.7 to 17.7°C for MAT and 1237 to 1414 mm for MAP. Rapoport’s Rule also helped to explain the elevational pattern of species richness in Yunnan, but those influences were not entirely uniform across all methods. These results suggested that the elevational pattern of orchid species richness in Yunnan is collectively shaped by several mechanisms related to geometric constraints, size of the land area, and environments. Because of the dominant role of climate in determining orchid richness, our findings may contribute to a better understanding of the potential effects of climate change on orchid diversity, and the development of conservation strategies for orchids.     

Elevational patterns of frog species richness and endemic richness in the Hengduan Mountains, China: geometric constraints, area and climate effects

We studied frog biodiversity along an elevational gradient in the Hengduan Mountains, China. Endemic and non-endemic elevational diversity patterns were examined individually. Competing hypotheses were also tested for these patterns. Species richness of total frogs, endemics and non-endemics peaked at mid-elevations. The peak in endemic species richness was at higher elevations than the maxima of total species richness. Endemic species richness followed the mid-domain model predictions, and showed a nonlinear relationship with temperature. Water and energy were the most important variables in explaining elevational patterns of non-endemic species richness. A suite of interacting climatic and geometric factors best explained total species richness patterns along the elevational gradient. We suggest that the mid-domain effect was an important factor to explain elevational richness patterns, especially in regions with high endemism.     

广东兰科植物多样性保育现状

为了使广东省的兰科植物及其遗传多样性得到有效的保育, 保存我国重要野生植物资源, 在2017-2019年间, 采用样线和样地相结合的调查手段、专家快速评估和野外调查相结合的评估技术以及Wilcoxon符号秩检验和Friedman检验的统计方法, 对广东省自然分布的兰科植物进行了全面的调查和濒危等级评估, 并对其在广东省自然保护区中的就地保育情况和全国植物园中的迁地保育情况进行了综合分析。结果表明, 广东省分布有兰科植物80属235种, 其中广东特有种20种; 广东兰科植物受威胁物种有186种, 其中极危11种、濒危114种、易危61种; 就地保育的兰科植物有111种, 迁地保育的兰科植物有156种, 就地和迁地共同保育的兰科植物有96种, 保育的有效程度较低; 另外, 就地、迁地、就地和迁地共同保育的兰科植物之间没有体现出明显的差异, 保育工作缺乏选择性和针对性。基于此, 我们建议广东兰科植物的保育工作应重视基础数据的收集和持续的野外监测、提高保育物种的数量、优化迁地保育物种的选择性和针对性、完善迁地保育和就地保育之间的协同性, 同时也应重视立法和公众教育, 并构建广东兰科植物保育的网络系统。     

Phylogenetic diversity,structure and diversification patterns of endemic plants along the elevational gradient in the Eastern Himalaya

ABSTRACT

Background: Several studies have documented the variation in species diversity patterns along elevational gradients in the Himalaya, but few have reported the evolutionary and biogeographic processes behind these patterns.

Aims: To understand whether evolutionary history and phylogeny have any role in structuring plant species communities along an elevational gradient in the Sikkim Himalaya.

Methods: We used data on endemic plant species occurrence from primary and secondary sources to construct family-level phylogenetic supertrees for different growth forms with the help of Phylomatic tool of Phylocom. These phylogenetic supertrees were used as a base for testing phylogenetic diversity (PD), niche conservatism, diversification time patterns and phylogenetic structure of various plant growth forms along an elevational gradient.

Results: PD was the highest at mid-elevations for all growth forms and PD had a significant positive correlation with endemic species richness. Species at mid-elevations were dominated by the ancestral/primitive taxa. There was phylogenetic clustering at higher elevations and phylogenetic overdispersion at lower and mid-elevations for the majority of the growth forms.

Conclusions: Time-for-speciation effect and niche conservatism along elevation (retention of niche-related ancestral elevational distribution over evolutionary time scale by species) together determine plant species diversity patterns in the Himalaya.     

高寒草甸群落植物多样性和初级生产力沿海拔梯度变化的研究

 对不同海拔梯度高寒草甸群落植物多样性和初级生产力关系的研究结果表明：1）不同海拔梯度上,中间海拔梯度群落植物多样性最高,即物种丰富度、均匀度和多样性最大;2）不同海拔梯度上,群落生产力水平和物种丰富度中等时,物种多样性最高;3）随着海拔的逐渐升高,地上生物量逐渐减少;4）地下生物量具有“V”字形季节变化规律,在牧草返青期和枯黄期地下生物量最大,7月最小,且地下生物量主要分布在0～10 cm的土层中。地下生物量垂直分布呈明显的倒金字塔特征。     

Patterns of an elevational gradient affecting moths across the South Korean mountains: effects of geometric constraints,plants, and climate

We investigated elevational richness patterns of three moth groups (Erebidae, Geometridae, and Noctuidae) along four elevational gradients located on one northern and three southern mountains in South Korea, as well as the effects of plants and climatic factors on the diversity patterns of moths. Moths were collected with an ultraviolet light trap at 32 sites from May through October, 2013. Plant species richness and mean temperatures for January and June were acquired. Observed and estimated moth species richness was calculated and the diversity patterns with null models were compared. Species richness along four elevational gradients peaked at mid-elevations, whereas deviations occurred at elevations below mid-peak in the southern mountains and elevations higher than mid-peak on the northern mountain. Species richness curves of three moth groups also peaked at mid-elevations throughout South Korea. However, the species richness curves for Erebidae were positively skewed, indicating that a preference for lowlands, whereas curves of the Geometridae were negatively skewed, indicating a preference for highlands. The mid-peak diversity pattern between plants and moths on the Korean mountains showed an elevational breadth that overlapped between 800 and 900 m. Multiple regression analysis revealed that plant species richness and January mean temperature significantly influenced moth species richness and abundance. The rapid increase in mean annual temperature in the Korean peninsula and the unimodal elevational gradients of moths across the country suggest that an uphill shift in peak optimum elevation and changes in the highest peak of the curve will occur in the future.     

Abundance variations within feeding guilds reveal ecological mechanisms behind avian species richness pattern along the elevational gradient of Mount Cameroon

Two distinct diversity patterns are observed along tropical elevations: (a) decreasing number of species toward high elevations and (b) a hump-shaped pattern with the peak at mid-elevations. As diversity is likely supported by ecological capacity of the environment, decomposition of the overall richness into ecological facets and considering number of individuals within them is crucial for the proper understanding of richness patterns. We examined abundances of different avian guilds along the forested part of the elevational gradient on Mt. Cameroon. We (a) compared richness and abundance elevational patterns, (b) assessed the effective contribution of multiple guilds to richness and abundance patterns, and (c) assessed to what extent observed abundances of guilds differed from those expected by chance. We sampled birds in 2011–2015 during the dry season at seven elevations (30 m, 350 m, 650 m, 1100 m, 1500 m, 1850 m, 2200 m a.s.l.). For each assemblage, we estimated proportions of species and individuals that use particular diets, foraging modes, and feeding strata. We found that a rather decreasing pattern of species richness turns into a hump-shaped one if we look at the total abundances, implying different mechanisms behind these patterns. The number of species and individuals thus do not seem to be directly related, contrary to “the more-individuals hypothesis.” Abundances of foliage gleaners at mid-elevations, nectarivores at high elevations, and frugivores at low elevations deviated from random expectations. Our results imply that parts of ecological space are filled separately by bird species and individuals along elevation of Mt. Cameroon.     

西双版纳片段化石灰岩森林附生兰科植物多样性研究

西双版纳石灰岩地区拥有丰富的兰科植物资源,但近年来随着橡胶树的大面积种植,使得该地区很多石灰岩森林呈片段化。为了解片段化石灰岩森林中附生兰科植物多样性状况以及片段化对附生兰科植物的影响,该研究选取了生境片段化的青岩寨和曼纳览,以及连续生境的绿石林和巴卡新寨等4个样地的29个样方进行多样性调查和对比研究。结果表明:4个石灰岩森林样共记录到附生兰科植物34属76种1 528株(丛)。通过对坡向、坡度、海拔、地形、郁闭度和森林类型等6个环境因子与附生兰科植物丰富度进行CCA分析,发现石灰岩地区附生兰科植物的分布主要受海拔和森林类型2个因素的影响,在海拔较高的青岩寨和巴卡新寨附生兰科植物物种丰富度高于低海拔的绿石林和曼纳览,而片段化对附生兰科植物的物种多样性和多度均无显著影响,这可能与片段化的历史较短有关。虽然目前来看生境片段化对石灰岩地区的附生兰科植物多样性无显著影响,但生境脆弱的石灰岩森林植被的保护对于兰科植物多样性保护则更具重要性。     

Elevational changes in vascular plants richness,diversity, and distribution pattern in Abune Yosef mountain range,Northern Ethiopia

The aim of this research is to investigate the patterns of vascular plant species richness,diversity,and distribution along an elevation gradient in the Abune Yosef mountain range,Ethiopia.Preferential systematic sampling was employed to collect vegetation and environmental data along the elevation gradient.We found that plant species richness declines monotonically from low to high elevations.Specifically,vascular plant species richness and diversity were lower in the Afroalpine grassland(high elevation)than in the Dry evergreen Afromontane forest and Ericaceous forest(low elevations).In contrast,endemic vascular plant richness was significantly higher in the Afroalpine grassland than in the Dry evergreen Afromontane forest and Ericaceous forest.Elevation showed a significant impact on the richness,diversity,and endemism of vascular plants.According to Sorensen's coefficient,the similarity between Dry evergreen Afromontane forest and Ericaceous forest vegetation types is higher(32%)than the similarity between Ericaceous forest and Afroalpine grassland(18%).Only 5%similarity was recorded between the Dry evergreen Afromontane forest and Afroalpine grassland.Growth forms showed different elevationai richness patterns.Trees and liana increased monotonically up to 3300 m.Shrub and herb richness patterns followed a hump-shaped and inverted hump-shaped pattern along the elevation gradient.The elevation patterns of vascular plant species richness,diversity,and growth form in the present study may be attributed to differences in management intensity,spatial heterogeneity,microclimatic variations,and anthropogenic disturbances.