(竹思)篣竹属及其近缘属叶表皮微形态特征

用扫描电镜和光学显微镜对(竹思)篣竹属(Schizostachyum Nees)7个种,及其近缘的梨竹属(Melocanna Trin.)、泡竹属(Pseudostachyum Munro)、薄竹属(Leptocanna Chia et H.L. Fung)各1个种以及空竹属(Cephalostachyum Munro)6个种的叶表皮微形态特征进行了观察.结果表明竹叶下表皮乳突的形状和在气孔器上及周围的排列式样在这5个属中各有不同,具有重要的分类学和系统学价值.同时我们的结果不支持广义的以属;而梨竹属、空竹属和泡竹属则被支持独立成属.另外,薄竹(Leptocanna chinensis)和红毛葸(竹思)篣竹(Schizostachyum sanguineum)的叶表皮微形态特征非常接近空竹属;香糯竹(Cephalostachyum pergracile)和金毛空竹(C.virgatum)的叶表皮微形态特征则十分接近(竹思)篣竹属.     

中国竹亚科空竹属的整理

我国竹亚科空竹属(Cephalostachyum Munro)植物发现于云南和西藏两省区,集中分布于云南。根据形态学性状,以及近年来叶片微形态和分子系统学研究结果,作者对我国空竹属物种进行了整理并编制了分种检索表。结果显示,我国空竹属现知有7种,云南均有分布:薄竹(Cephalostachyum chinense(Rendle)D.Z.Li et H.Q.Yang)、空竹(C.fuchsianum Gamble et Hook.f.)、独龙江空竹(C.mannii(Gamble)Stapleton et D.Z.Li)、小空竹(C.pallidum Munro)、屏边空竹(C.pingbianense(Hsueh et Y.M.Yang ex Yi et al.)D.Z.Li et H.Q.Yang)、红毛空竹(C.sanguineum(W.P.Zhang)D.Z.LiH.Q.Yang)、真麻竹(C.scandens Bor)。薄竹属(Leptocanna Chia et H.L.Fung)和毒空竹(C.virulentum Y.M.Yang et F.Du)分别为空竹属和空竹的异名。另一方面,原置于空竹属的香糯竹(C.pergracile Munro)和金毛空竹(C.virgatum(Munro)Kurz)的形态特征则更接近狭义的竹属(Schizostachyum Nees),应该处理为狭义竹属成员。本研究对空竹属今后的修订工作有重要意义。     

中国(竹思)簩竹属(Schizostachyum Nees)的研究及其它

对国产(竹思)(竹劳)竹属(Schizostachyum Nees)进行了系统整理。对属的范围进行了修订,认为本属应包括乔草竹属(Dendrochloa Parkinson),薄竹属(Leptocanna Chia et H.L.Fung),长穗竹属(Teinostachyum Munro)和李海竹属(Neohouzeaua A.Camus),并首次建立了本属的属下分类系统。此外,对二种竹子起了新名:甲竹(Bambusa austro-sinensis Xia)和毛环单竹(B.yunnanensis Xia);作了4个新组合,即Schizostachyum coradatum(Wen et Dai)Xia, S.dumetorum(Hance)Munro var.xinwuense(Wen et J. Y Chin)Xia, Bambusa glaucescens glaucescens(Willd)Sieb. ex Munro var.annulata(W.T.Lin et Z.J.Feng)Xia和B.glaucescens (Willd)Sieb.ex Munro var.pubivagina(W.T.Lin et Z.J.Feng)Xia.     

中国(竹思)簩竹属(Schizostachyum Nees)的研究及其它

对国产(竹思)(竹劳)竹属(Schizostachyum Nees)进行了系统整理。对属的范围进行了修订,认为本属应包括乔草竹属(Dendrochloa Parkinson),薄竹属(Leptocanna Chia et H.L.Fung),长穗竹属(Teinostachyum Munro)和李海竹属(Neohouzeaua A.Camus),并首次建立了本属的属下分类系统。此外,对二种竹子起了新名:甲竹(Bambusa austro-sinensis Xia)和毛环单竹(B.yunnanensis Xia);作了4个新组合,即Schizostachyum coradatum(Wen et Dai)Xia, S.dumetorum(Hance)Munro var.xinwuense(Wen et J. Y Chin)Xia, Bambusa glaucescens glaucescens(Willd)Sieb. ex Munro var.annulata(W.T.Lin et Z.J.Feng)Xia和B.glaucescens (Willd)Sieb.ex Munro var.pubivagina(W.T....     

Chi竹属和牡竹属（竹亚科）叶表皮微形态特征

利用光学显微镜和扫描电镜,对Chi竹属21个种（包括Chi竹亚属5个种、孝顺竹亚属2种、单竹亚属4种绿竹亚属9种和亚属未定1种）以及牡竹属5种叶表皮微形态特征进行了观察和研究。所得结果表明气孔保卫细胞上乳突的数目和分布存在一定的规律,并具有一定的分类价值。绿竹亚属的叶表皮微形态特征界于Chi竹属和牡竹属之间,但更接近于Chi竹属。因此认为在分类处理上,将绿竹亚属置于Chi竹属比单独成立绿竹属更为合理。     

箣竹属和牡竹属(竹亚科)叶表皮微形态特征

利用光学显微镜和扫描电镜,对箣竹属21个种(包括箣竹亚属5个种、孝顺竹亚属2种、单竹亚属4种、绿竹亚属9种和亚属未定1种)以及牡竹属5种叶表皮微形态特征进行了观察和研究。所得结果表明气孔保卫细胞上乳突的数目和分布存在一定的规律,并具有一定的分类价值。绿竹亚属的叶表皮微形态特征界于箣竹属和牡竹属之间,但更接近于箣竹属。因此认为在分类处理上,将绿竹亚属置于箣竹属比单独成立绿竹属更为合理。     

应用叶表皮结构讨论槽稃草属Euthryptochloa Cope(Gramineae)的归属问题

比较研究单种属,槽稃草属Euthryptochloa Cope与显子草属Phaenosperma Munro的叶表皮微形态,并结合外部形态特征说明槽稃草属与显子草属无论在叶表皮微形态或外部形态上都无区别。根据国际植物命名法规的优先律,槽稃草属不能独立成属,应归入显子草属。     

井冈寒竹属叶表皮微形态特征及其分类学意义

井冈寒竹属(又名短枝竹属,Gelidocalamus Wen)是1982年温太辉以产于井冈山区的井冈寒竹Gelidocalamus stellatus Wen为模式种建立的新属。自井冈寒竹属建立以来,陆续有新的分类群发表,或将一些分类群归并到本属中,但迄今尚未开展系统研究。为了发现更多的井冈寒竹属分类学证据,本研究利用扫描电镜对井冈寒竹属9个分类群的叶表皮微形态特征进行了观察。结果表明:9个分类群竹叶下表皮的长细胞、微毛、大毛的形态和分布较为一致,但乳突、刺毛、气孔器和短细胞呈现较丰富的微形态差异。尤其是乳突的形状、气孔器及其周围的乳突排列式样最为丰富,具有重要的分类学价值,可以作为属内种的划分依据。     

《中国植物志》(英文版)竹亚科青篱竹属和新小竹属的修订

基于最近的分子系统学研究、近期发表的新类群以及国际植物命名法规的相关条款,对《中国植物志》 （英文版）竹亚科的两个属进行了修订。中国是否有青篱竹属 (Arundinaria)的分布一直是个争论不休的问题。分子系统学不支持广义青篱竹属,因此原置于青篱竹属下的4个种应恢复到巴山木竹属 (Bashania) 中。包括近期发表的新类群在内,巴山木竹属在中国共有10种。西藏新小竹 (Neomicrocalamus microphyllus)是一个没有合格发表的裸名,其正确名称应为新小竹 (N.prainii),而云南新小竹（N.yunnanensis）则可能是梨籐竹属的成员。     

巴西竹类生物多样性

巴西是全球生物多样性最丰富的地区,其竹类多样性也极为丰富。结合现存资料及野外调查,对巴西全境的竹类分布格局进行了讨论。巴西全国有原生竹亚科植物256种(含2亚种及3变种),北部地区草本竹类(莪莉竹族)最丰富,有61种,而东南部地区木本竹类(箣竹族)最丰富,有96种。偏穗竹属(Merostachys)(43种)和丘斯夸竹属(Chusquea)(45种)是最常见的属,并是最具潜在经济利用的竹类。属种的特有性分别高达32.4%和68.8%。特有属有11个,分别为莪莉竹族的双药莪利草竹属(Diandrolyra)、独焰草竹属(Eremitis)、小百瑞草竹属(Parianella)、赖茨草竹属(Reitzia)、苏克蕾草竹属(Sucrea)和箣竹族的南美梨藤竹属(Alvimia)、离枝竹属(Apoclada)、密穗竹属(Athroostachys)、卡姆巴珠瓦竹属(Cambajuva)、菲尔盖拉斯竹属(Filgueirasia)、无枝竹属(Glaziophyton)。     

Micromorphological Study on Leaf Epidermis of Schizostachyum and Its Allies ( Poaceae: Bambusoideae)

Micromorphological characters of leaf epidermis of 16 published species from Schizostachy um Nees and its allies ( including Melocanna Trin. , Pseudostachyum Munro, Leptocanna Chia et H. L. Fung and Cephalostachyum Munro) were examined by both light and scanning electron microscopies. The results indicated that the papillae forms and distributional patterns on the abaxial surface were taxonomically informative at the generic level. The characters of papillae did not support Schiz ostachyum sensu lato; meanwhile, it was supported that Melocanna, Pseudostachyum and Cep halostachyum were better to be treated as separate genera respectively. Furthermore, it seemed to be more appropriate to place Leptocanna chinensis and Schizostachyum sanguineum in Cep halostachyum; and to place Cephalostachyum pergr acile and C1 virgatum in Schizostachy um .     

A molecular phylogenetic and fruit evolutionary analysis of the major groups of the paleotropical woody bamboos (Gramineae: Bambusoideae) based on nuclear ITS, GBSSI gene and plastid trnL-F DNA sequences

This study presented the first molecular phylogenetic analysis of the major clades of woody bamboos of the Old World tropics based on nuclear and chloroplast sequences (ITS, GBSSI and trnL-F). Sequence data from 53 species, representing 17 paleotropical woody bamboo genera, were analyzed using the maximum parsimony and Bayesian inference methods. All examined ingroup taxa were clustered into two clades, i.e., the Bambusinae+Dinochloa clade and the Melocanninae clade. The former clade included Bambusa, Bonia, Dendrocalamus, Dendrocalamopsis, Dinochloa, Gigantochloa, Molecalamus, Neomicrocalamus, Neosinocalamus, Oxytenanthera s. str. (sensu stricto), Racemobambos and Thyrsostachys. The Melocanninae clade consisted of Cephalostachyum, Leptocanna (better treated as part of Cephalostachyum), Melocanna, Pseudostachyum and Schizostachyum s. str. The subtribe Racemobambosinae and tribes Dendrocalameae and Oxytenanthereae were not supported and may be better placed in subtribe Bambusinae. The ovary characters seemed to be good criteria to distinguish these two clades. The reconstruction of ancestral fruit characters indicated that the bacoid caryopsis, namely, fleshy or berry-like fruits, was found to be scattered in three lineages of the examined paleotropical woody bamboos. Fruit characters are thus not reliable indicators of phylogeny and bacoid caryopsis likely represents a specialization for particular ecological conditions.     

Comparative morphology of the foliage leaf epidermis, with emphasis on papillae characters, in key taxa of woody bamboos of the Asian tropics (Poaceae: Bambusoideae)

The foliage leaf epidermis of 35 species representing 12 key genera of woody bamboos of the Asian tropics was investigated using light and scanning electron microscopy. The results indicated that papillae forms and distributional patterns around the stomatal apparatus of the abaxial foliage leaf epidermis were usually constant and were of great taxonomic significance at the specific and generic levels. However, papillae characters were not suitable for dividing subtribes within woody bamboos of the Asian tropics. On the basis of papillae characters, Schizostachyum s.s. and Cephalostachyum were confirmed, but their delimitations should be modified. The transfer of Leptocanna chinensis and Schizostachyum sanguineum into Cephalostachyum was supported, and Cephalostachyum virgatum and C .  pergracile were confirmed to be members of Schizostachyum s.s. The subtribe Racemobambosinae did not obtain support and Racemobambos appeared to be better placed in subtribe Bambusinae. Neomicrocalamus was supported as a close relative and better treated as a synonym of Racemobambos . Gigantochloa was closely related to Dendrocalamus .  © 2008 The Linnean Society of London, Botanical Journal of the Linnean Society , 2008, 156 , 411–423.     

Revision on Cephalostachyum Munro (Poaceae: Bambusoideae) in China

Species of Cephalostachyum Munro (Poaceae: Bambusoideae) from China are distributed in Yunnan and Tibet, mainly in Yunnan. In this paper, we discussed species of Cephalostachyum and compiled a key to species from China, based on recent studies on micromorphological characters of leaf epidermis and molecular phylogenetics of paleotropical woody bamboos. There is a total of seven species of Cephalostachyum distributed in China, all in Yunnan: Cchinense (Rendle) DZ. Li et HQ. Yang, Cfuchsianum Gamble et Hook. f., Cmannii (Gamble) Stapleton et DZ. Li, Cpallidum Munro, Cpingbianense (Hsueh et YM. Yang ex Yi et al.) DZ. Li et HQ. Yang, Csanguineum (WP. Zhang) DZ. Li et HQ. Yang and Cscandens Bor. Leptocanna Chia et HL. Fung and Cvirulentum YM Yang et F. Du are synonyms of Cephalostachyum Munro and Cfuchsianum Gamble et Hook. f. respectively. On the other hand, Cpergracile Munro and Cvirgatum (Munro) Kurz are morphologically closer to Schizostachyum Nees than to Cephalostachyum, and they should be treated as members of Schizostachyum. This paper is of significance to a worldwide revision of Cephalostachyum.     

The Morphology of Fruits and Starches in Bamboos,and Its Relation to Systematic Position

In this article, 30 speceis of bamboos, including 19 genera in 5 tribes, were collected and the morphology of fruits and starches of them was studied. The results are as follows. I. The morphology of fruits is important in studies of systematic position in bamboos. According to the systems of W. Munro and G. Bentham whether the pericarp is adhesive to or free from the seed coat may be taken as a basis of classification. It is also confirmed in this article. It is found in this work that all taxa with a binding pericarp and seed coat are of caryopsis that also has a ventral suture and hilum, while all others with a separated pericarp and seed coat are of bacca or nut, which has no ventral suture and hilum. The former has a hard and thin pericarp and rich endosperm, while the latter has a fleshy and thick pericarp and no endosperm. These characteristics form a basis of classification of major groups. II. In 1907, Brandis found that no any endosperm in matured fruit of Dinochloa, Melocalamus, Melocanna and Ochlandra. It has been proved by Stapf in at least one genus. We found that the baccae of Qiongzhuea, Melocanna, Ferrocalamus and Chimonobambusa Subg. Oerocalama were empty, with no endosperm. This may be a common character of the bacca. We believe, therefore, that the systematic position of Qiongzhuea, Ferrocalamus and Chimonobambusa Subg. Oreocalama is close to Melocanneae. III. Starch grains of bamboo fruits are complex in structure. They are round or ellipsoidal, consisting of 3-22 polyhedral or apple-like small grains. The morphology of starch grains is not so important as fruit in bamboo classification, but some characteristics are of a high value in the identification of genera and species, when they are combined with other features. In Cephalostachyum, the starch grain is very big, with 20-40 μm in diam, and the starch small grain is polyhedral or apple-like with 7.5-22.5 μm in diam, while in Dendrocalamus, the starch grain is small, with 10-28.9 μm in diam. and the starch small grain is only polyhedral, with 3-11.9 μm in diam. The morphology and size of the starch grain and starch small grain are also different in Melocanna and Chimonobambusa Subg. Oreocalama. IV. W. Munro’s system divided Bambuseae into three major groups according to the morphology of flower and fruit. Because the material was not sufficient at that time, the system wrongly put Cephalostachyum, Dendrocalamus into the group Bacciferea. Now it is found that both Cephalostachyum and Dendrocalamus have a nut. Later G. Bentham found this problem and divided the Bambuseae into four subtribes, treating Dendrocalamus as a separate subtribe, Dendrocalamae, and putting the bacca group into another subtribe, Melocannae. It is better, but it also has some shortcomings. Hackel, Gamble, E. G. Camus, A. Camus and Keng Pojie all accepted the view of Bentham, placing Dendrocalamus and Melocanna into different subtribes or tribes.     

Flowering in a bamboo, Melocanna baccifera (Bambusoideae: Poaceae)

Melocanna baccifera (Roxburgh) Kurtz ex Skeels, a species of bamboo introduced to Sri Lanka from India, flowered and set fruit during 2001–2002. Culms that flowered and set fruit died. The incidence of flowering is significant in that flowering took place close to the predicted mast flowering in 2007. At the onset of flowering, inflorescences were predominantly staminate. But later in 2002, bisexual and pistilate flowers also developed leading to fruiting. Both protandry and protogyny were observed in the bisexual florets. Floral characters indicated that the species was mainly out-crossing. Although anther dehiscence released pollen and stigmas were exerted, pollination of stigmas was inefficient. The few stigmas that were naturally pollinated showed limited pollen tube growth. However, fruit set took place. Fruits were very rarely seen to germinate naturally. Many fruits were devoid of an embryo, indicating that parthenocarpy and/or apomixis may have taken place. Excised embryos germinated in vitro .  © 2003 The Linnean Society of London, Botanical Journal of the Linnean Society , 2003, 143 , 287–291.     

Variability in epidermal characters of <Emphasis Type="Italic">Ginkgo tzagajanica</Emphasis> Samylina (Ginkgoales) from the Paleocene of the Tsagayan formation (Amur Region) and the taxonomy of tertiary species of <Emphasis Type="Italic">Ginkgo</Emphasis>

The study of new material on Ginkgo tzagajanica Samylina and earlier published data has shown that the epidermal morphology of the species in highly variable; several new characters were found. Occurrence of different states of characters was analyzed, their frequency distribution was revealed, and typical and rare states of characters within variation rows were determined. The specific diagnosis of G. tzagajanica was emended. The most variable characters are degree of undulation of the anticlinal walls of cells of abaxial and adaxial epidermises (the latter is more variable) and degree of development and number of papillae on ordinary epidermal cells. Less variable characters are degree of development of papillae on subsidiary cells of stomata and how the papillae cover the stomatal apertures. These are characters that bear maximal diagnostic significance for Cenozoic Ginkgo.