L’industrie lithique des gisements du Pléistocène inférieur de Barranco León et Fuente Nueva 3 à Orce, Grenade, Espagne

The archaeological localities of Barranco León and Fuente Nueva 3 in Orce, Grenade, in actuality have the oldest lithic artifacts in Europe; they are the oldest evidence of human occupation in the east of Europe. When compared the results from faunal analysis and magnetostratigraphical study, notably through the presence of Allophaiomys lavocati allow for a date of around 1.3 Ma. The raw material used for developing these lithic industries are, basically limestone or siliceous rocks, these raw materials selected were smoothed down in situ. The techniques used are knapping at raised with a hammerstone, and we can find all the different elements of the “chaine opératoire”. Both localities are characterised by the predominance of flakes and debris, cores, and some knapped cobbles sowing mediocre aspect. The technical and typological features of both lithic assemblages have an opportunist character but they are relatively sophisticated too.     

Lithostratigraphie et biostratigraphie des formations paléocènes et éocènes entre Bayonne et Pau (SW France)

The lithostratigraphy and biostratigraphy of the Paleocene and Eocene formations have been reconsidered on the basis of 18 dated boreholes that intersect the Basque Country, Peyrehorade, Sainte-Suzanne and Sauvelade allochthonous tectonic units, as well as the autochthonous units of the Pey-Saint-Lon anticline, the Arzacq Basin and the North Pyrenean Trough. Correlation of the formations identified in the boreholes and at outcrop has made it possible to outline the sedimentary evolution of this region during the Early Paleogene and to date the various stages of Pyrenean compression on the basis of sedimentary gaps and syntectonic detrital input. The Lasseube Formation is present over the entire study area. Sedimentation becomes differentiated during the Late Selandian-Early Ypresian period. In the North Pyrenean Trough and in the Peyrehorade, Sainte-Suzanne and Sauvelade units, the detrital-clayey Pont-Labau Formation is well developed and all its members are represented (Guillempau, Latapy, Néez, Libaros and Artigueloutan; zones upper P 3b-P 6b; upper NP 5-NP 11). A sedimentary gap showing erosion, and involving zones upper P 3b-upper P 4a and upper NP 5-NP 7, characterises the Basque units, the Pey-Saint-Lon anticline and the western part of the Arzacq Basin. Sedimentation recommences in zone P 8 with middle-bathyal deposits of the Cassoua Member and then continues with the Artigueloutan Member (transition facies towards the Louts Formation) or limestone resembling that of the Meilhan Formation. The Gan Formation shows an east-west polarity. In the east, between the Sainte-Suzanne Unit and the North Pyrenean Trough, the formation is argillaceous with a dominant neritic character, apart from a circalittoral to epibathyal passage in its lower part (zone P 7) containing numerous planktonic foraminifera. The formation is of a detrital and carbonate nature in the west (Sainte-Marie-de-Gosse 1). The bathyal Donzacq Formation appears to transgress a previously deformed surface. Two domains existed during the Lutetian: a continental to neritic domain in the south and a bathyal domain in the north (the Saint-Geours-d'Auribat and Miretrain formations) separated by a variably thick transition zone that is affected by thrust fronts. Two new formations - the Orthez and Sorde-l'Abbaye formations - are proposed, and the Marly limestone formation of Saint-Barthélémy (Douvillé, 1905) is redefined. The Orthez Formation (Sauvelade Unit) is composed of marl and continental conglomerate with neritic carbonate interbedding and belongs to the sub-zones NP 15a and b. The Sorde-l'Abbaye Formation (Peyrehorade Unit and the Basque units p. p.) is composed of neritic carbonate sediments (Lutetian-Priabonian). The Saint-Barthélémy Formation is made up of syntectonic conglomerate, turbiditic limestone and pelagic marl in the central part of the Basque units (NP 15a-NP 16). The Bartonian and Priabonian stages are characterised by carbonate sediments in the Arzacq Basin (Siest Formation). Its base is diachronous: Upper Lutetian (zone SB 16) in the east and Late Bartonian grading to Priabonian in the west (zones SB 17/18 and SB 19). The sediments are argillaceous (Côte des Basques Formation) along the Atlantic coast, in the central part of the Basque units and in the Bayonne-Cauneille syncline. Major thickness variations reflect tectonic phases at the top of the Lasseube Formation in the north (erosion pre NP 8), the Néez Member (NP 7/NP 8 boundary), the Artigueloutan Member (NP 11/NP 12 boundary), and the Gan Formation (P 8/P 9 boundary). The syntectonic conglomerate and breccia indicate the development of thrust fronts in the Libaros Member (NP 8), at the base of the Pillardou and Artigueloutan members (NP 9b, c-NP 10), in the Saint-Geours-d'Auribat Formation (NP 14b) and in the Orthez and Saint-Barthélémy formations (between zones NP 15a and NP 16).     

Les foraminifères du Paléocène et de l'Éocène basal du sillon nord-pyrénéen (Aquitaine, France)

This paper presents for the first time the inventory of the Paleocene and Lower Eocene foraminifers located in the North Pyrenean trough, between the Atlantic Ocean and the neighbourhood of the town Pau. They have been studied from three outcrops. The Bidart Beach section shows the Lasseube Formation from the Cretaceous/Paleogene boundary to the base of the P 3a zone. The Loubieng Quarry section, near Orthez, represents the upper P 3a zone and the lower P 3b zone as well as the Lasseube Formation / Pont Labau Formation boundary. The interval between the upper part of the P 3b zone and the upperest part of the P 5 zone crops out along the Gan - Rébénacq road with a hiatus located at the Paleocene / Eocene boundary, the whole interval belonging to the Pont Labau Formation. 394 taxons of foraminifers are present in this formations: 349 benthic and 45 planktonic species. The Velasco type benthic foraminifers show a middle bathyal depositional environment, with a paleobathymetry included between 500-600 m and 1000 m: Nuttallides truempyi, Osangularia velascoensis, Bulimina trinitatensis. The Midway type species which were transported by the turbidite currents from the lower to middle neritic environments are frequent as well as the Cretaceous reworked species. The species number is low: 29 in the iridium layer of the Cretaceous/Paleogene boundary (P 0 zone). Fauna grows rich quickly in the Pα zone reaching 129 species. The diversity grows up progressively from P α to the P 4a zone (NP 8). The disappearances are rare until his horizon, but their number is bigger than the number of appearances from the P 4b zone. It reaches their maximum between the P 4c zone and the P 5 zone. It shows that the decline of the Paleocene fauna begins around 2 million years before the thermal event of the Paleocene / Eocene boundary in the Aquitaine sections. The disappearances stay important in the Lower Eocene - Ypresian, but the appearance of lots of Eocene species show that the fauna renewal is located in this stage. The Cretaceous taxa dominate in the Paleocene benthonic fauna. The appearance or the disappearance of some species has a stratigraphic value in the Aquitaine region: the disappearance of Coryphostoma incrassata in the P 1b zone, the appearances of Plectina dalmatina, Elongobula grata (P α); Bulimina tuxpamensis (P 2); Tritaxilina cubensis, Thalmannita madrugaensis (P 3a); Svenia bulbosa (P 3b); Discorbis perovalis (P 4a/NP 7); Elongobula pulchra, E. pupa, Asterigerina bartoniana, Neorotalia gr. tuberculata (P 4a/NP 8); Bigenerina pannonica, Pentellina pseudosaxorum (P 5/NP 9-10).     

Les faunes d’ammonites du Pliensbachien inférieur des séries dauphinoises de la Durance (Jurassique inférieur, Barrage de Serre-Ponçon, Hautes-Alpes, France)

The Liassic ammonite faunas and especially those of the Early Pliensbachian are of major importance for the understanding of the stratigraphic framework of the “Dauphiné” zone of the southern subalpine ranges (“dôme de Remollon”). Until now, only listed by the authors, these faunas are, for the first time, described and illustrated in the present paper. With about twenty species, these faunas cover the whole of the Early Pliensbachian. There are poorly diversified and of clearly Euroboreal affinities. Nearly all of the species are abundant and common in most of the NW European fossiliferous localities. Despite of a paleogeographical location fairly close to the Alpine zones (Southern Alps and Austroalpine units) with rather diversified faunas of mainly Mediterranean affinities, none Mediterranean taxa are present in the studied area. Unfortunately, the current stratigraphic and paleogeographic knowledge do not allow to ascertain if this reflects a separation due to paleoenvironmental or true paleogeographic barriers. In this latter case, the general geologic implications would be remarkable.     

Biochronologie mammalienne du Quaternaire du Maroc atlantique, dans son cadre régional

Recent researches conducted in Morocco in the frame of the “Programme Casablanca” provide a firm ground to set up a biochronology of the Pliocene and Pleistocene based upon the succession of Mammalian faunas, thanks to the occurrence of well-dated landmarks, and despite very scarce absolute datations. The site of Lissasfa, dated to the Miocene-Pliocene boundary by rodents, testifies to important exchanges with Europe. That of Ahl al Oughlam yielded more than 100 vertebrate species. The evolutionary stages of its rodents, carnivores, bovids, Kolpochoerus, as well as the absence of Equus, allow us to refer it to the late Pliocene, at ca. 2.5 Ma. The Thomas and Oulad Hamida quarries in Casablanca cut through a unique succession of marine and continental levels going back to the lower Pleistocene, yielding the earliest evidence of human presence in Morocco. Later stages of this human occupation are documented by the Middle Pleistocene levels of these quarries, and by several other sites, most of which are also located in the Rabat-Casablanca region, thus confirming the importance of this coastal area in human history. However, the first chapter of this history in North-western Africa, at present illustrated by the sites of Aïn Hanech and Mansourah in Algeria, remains poorly documented in Morocco.     

Les stratégies de débitage dans les industries lithiques archaïques des premiers habitants de l’Europe

Barranco León and Fuente Nueva 3 sites at Orce in the south of Spain, and Vallonnet cave in the south of France, are some of the oldest sites in Europe, testimonies of the first settlements of these territories. The Orce sites, dated around 1.3-1.2 Ma, have given away numerous lithic artifacts principally issued from an important débitage activity. The beginning of flake production processes is few represented into these two consummation sites. In return, full débitage flakes are in high proportion, with generally short dimensions, in consequence of which, in some case an intense core reduction. Two percussion methods are identified: with direct percussion or on anvil. Sometimes, there was débitage on flake, taking advantage of the inferior face natural convexity of the core-flake, permitting the maximal exploitation and the economy of the best raw materials. The bipolar percussion on anvil permitted the maximal exploitation of the short dimensions supports, often in naturally cubic or sub-cubic form. A lot of the producted flakes, sometimes called “pièces esquillées”, show some scales on the distal and/or proximal transversal edges, issued from this bipolar knapping method. In the Vallonnet cave, where the occupation levels are dated around 1 Ma, the lithic industry show a lot of percussion tools and hammerstone flakes, but also some core reduction flakes and some very rares cores, especially in flint. This good raw material was chosen for the débitage, and the flakes show on superior face several anterior removals, and a butt which could reveal a striking platform preparation. At the gates of Europe, in Georgia, Dmanisi site give away an abundant lithic industry associated with faunal and human remains, dated around 1.81 to 1.77 Ma. Amid the lithic assemblages, cores are rather frequent and are characterized by a poor degree of exploitation, most often unifacial, and high frequency of cortical striking platforms. Raw materials, essentially volcanic cobbles, were from local origin, and the knapped supports were often chosen for their angular morphology adequate to begin the débitage in easier way, débitage which was essentially by direct percussion.     

内蒙古二连盆地古近系研究回顾及存在问题

简要回顾了内蒙古二连盆地古近系研究的历史,介绍了最新的研究进展,同时对主要地点的位置以及地层发育情况进行了讨论,并针对不同地点或地区分析了二连盆地古近系划分对比中存在的问题,对化石产地的地名和岩石地层单位名称的使用提出了建议。     

New Oldowan localities at high level within Kilombe Caldera,Kenya

New occurrences of early artefacts ascribed to the Oldowan tradition come from localities at high level within the caldera of the extinct Kilombe volcano, located in the central rift valley of Kenya. The trachyte cone and caldera of Kilombe volcano formed at ca. 2.5 Ma, and the record of >130 m of sediment-fill indicates that the caldera subsequently held a lake for long periods during the Early Pleistocene. The Oldowan artefact localities, dated by ⁴⁰Ar/³⁹Ar and palaeomagnetism to ～1.78 Ma, lie east of the centre of the caldera, on the west side of an ancient small lake, which later drained away as a gorge formed on the east side of the mountain. The artefacts are dominantly made of Kilombe trachyte, and are associated with a fauna of large animals including Hippopotamus gorgops. These are the first Oldowan localities to be discovered in a new area of the Kenyan rift valley in the last thirty years, and their presence at high level in rugged landscape indicates that the associated hominins were exploiting a full range of environments.     

Live (Rose Bengal stained) and dead benthic foraminifera from the oxygen minimum zone of the Pakistan continental margin (Arabian Sea)

Live (Rose Bengal stained) and dead benthic foraminiferal communities (hard-shelled species only) from the Pakistan continental margin oxygen minimum zone (OMZ) have been studied in order to determine the relation between faunal composition and the oxygenation of bottom waters. Samples were taken from 136 m to 1870 m water depth during the intermonsoon season of 2003 (March–April). Live foraminiferal densities show a clear maximum in the first half centimetre of the sediment only few specimens are found down to 4 cm depth. The faunas exhibit a clear zonation across the Pakistan margin OMZ. Down to 500 m water depth, Uvigerina ex gr. U. semiornata and Bolivina aff. B. dilatata dominate the assemblages. These taxa are largely restricted to the upper cm of the sediment. They are adapted to the very low bottom-water oxygen values (≈ 0.1 ml/l in the OMZ core) and the extremely high input of organic carbon on the upper continental slope. The lower part of the OMZ is characterised by cosmopolitan faunas, containing also some taxa that in other areas have been described in deep infaunal microhabitats. The contrast between faunas typical for the upper part of the OMZ, and cosmopolitan faunas in the lower part of the OMZ, may be explained by a difference in the stability of dysoxic conditions over geological time periods. The core of the OMZ has been characterised by prolonged periods of stable, strongly dysoxic conditions. The lower part of the OMZ, on the contrary, has been much more variable over time-scales of 1000s and 10,000 years because of changes in surface productivity and a fluctuating intensity of NADW circulation. We suggest that, as a consequence, well-adapted, shallow infaunal taxa occupy the upper part of the OMZ, whereas in the lower part of the OMZ, cosmopolitan deep infaunal taxa have repeatedly colonised these more intermittent low oxygen environments.     

Earliest occurrence of Puma pardoides (Owen, 1846) (Carnivora,Felidae) at the Plio/Pleistocene transition in western Europe: New evidence from the Middle Villafranchian assemblage of Montopoli,Italy

The puma-like cat Puma pardoides (Owen, 1846) is reported from several Eurasian localities dated to the Late Pliocene–Early Pleistocene, but its Italian fossil record is really poor; in fact, the only possible occurrence comprises some fragmentary postcranial material from the Late Villafranchian locality of Pirro Nord (1.6–1.3 Ma). In the present paper, we describe an isolated left upper carnassial of a medium-sized felid belonging to the collection of fossil mammals from Montopoli (Tuscany, Italy), the type locality of the Middle Villafranchian Faunal Unit (∼2.6 Ma). All the morphological and morphometric characters of the tooth are consistent with an attribution to Puma pardoides (Owen, 1846), which comes out as the second felid species from Montopoli together with Acinonyx pardinensis (Croizet et Jobert, 1828) and is, possibly with the specimens from Perrier-Étouaries, the earliest occurrence in western Europe.     

Les foraminifères du Burdigalien moyen à supérieur de la région sud-aquitaine (golfe de Saubrigues, SW France)

Many exposures assigned to the Miocene formation of “Marnes de Saubrigues” (southwestern Aquitaine Basin) have been studied. Their revised stratigraphical allocation has made it possible to recognize mainly Middle to Late Burdigalian age deposits which have been widely sampled. They have been correlated with planktonic zones N6 pars-N7 and NN3-NN4 and locally dated 17.7 Ma with Sr isotopes. It is noteworthy that this chronological interval is only known in the southern Basin. An inventory of the rather abundant microplankton is provided, including series crossed by drillings in the Gulf of Biscay. These deposits yield a large diversity of benthic foraminifera, nearly 400 taxa, which can be considered as a reference fauna for western Europe Burdigalian. From a geographic point of view, it has been possible to subdivise the exposures into groups according to their bathymetry. Let us point out the first discovery of Borelis in the Burdigalian of France and in the northeastern Atlantic Miocene as well. Very rare specimens of Cycloclypeus are also present in the Aquitaine Basin. Among smaller foraminifera, the occurrence of Rosalina aguayoi, Pavonitina styriaca, the last appearance of Falsocibicides aquitanicus and the persistance of the genus Almaena must be emphasized. Several assemblages supply diatoms, rare radiolaria and sometimes abundant sponge spicules; the presence of these organisms, seldom reported from the Aquitaine Tertiary, might be linked to upwelling-type currents. Bachmayerella (inc. sed.) is uncommon and cited for the first time in the Burdigalian. A few taxonomical remarks and short notes are given for some species of foraminifera, and palaeoecological details as well. Numerous taxa are represented and comparisons with the foraminifera faunas mainly from the Lower-Middle Miocene of Western Mediterranean and Paratethys are sketched.     

Rongeurs et lagomorphes du Miocène inférieur de Béon 2 (MN4, Montréal-du-Gers,SW France)

About 200 micromammal isolated teeth (rodents, lagomorphs) originating from the middle Orleanian locality of Béon 2 (Montréal-du-Gers, SW France) are described. The rodent fauna is dominated by myomorphs (7 species), including the cricetid Democricetodon aff. hispanicus Fahlbusch, the melissiodontid Melissiodon sp., the glirids Peridyromys murinus (Pomel), Pseudodryomys aff. ibericus, Pseudodryomys aff. simplicidens, and Glirudinus modestus (Dehm), and the eomyid Ligerimys aff. florancei. Sciuromorphs are represented only by Heteroxerus rubricati Crusafont, Villalta and Truyols. Within lagomorphs, 2 dental morphs referred to the ochotonid Prolagus Pomel are identified in Béon 2. They correspond to Prolagus oeningensis (König) and P. aff. vasconiensis. On a biostratigraphical point of view, this study confirms the location of Béon 2 at the early part of the MN4; this locality is older than other regional sites such as Pellecahus, Béon 1, La Romieu, and Bézian (MN4b), and it is coeval to Artenay, in the Loire basin. In particular, glirids and lagomorphs from Béon 2 testify to close relationships with micromammal localities referred to the MN3 biozone.     

On the status of Isolophodon Roth, 1903 (Mammalia,Astrapotheria) and other little-known Paleogene astrapotheres from central Patagonia

Most of the 16 currently recognized astrapothere genera are well known through numerous specimens preserving at least almost complete dentition. One of the exceptions is the enigmatic genus Isolophodon Roth, 1903, based on very scant and fragmentary materials from Paleogene levels of central Patagonia. This taxon was ruled out from almost all taxonomic lists, although its validity has not been discussed by subsequent authors. We herein re-describe and discuss the taxonomic status of the species of Isolophodon. The type species, I. cingulosus Roth, 1903, is characterized by having lower cheek teeth with a much reduced hypoflexid, resembling derived uruguaytheriines, but lower-crowned and with three lower premolars, as in the species of Astraponotus Ameghino, 1901. This is the only astrapothere nominal species attributable to the Tinguirirican South American Land Mammal Age (SALMA, Early Oligocene). Isolophodon aplanatus Roth, 1903 (Casamayoran and Mustersan SALMAs, middle Late Eocene) has proportionally more elongated lower molars and a less developed paraflexid than the type species. Isolophodon would represent an early diverging lineage of astrapotheriids, in which some dental features evolved convergently with the more derived uruguaytheriines. Additionally, we describe other fragmentary but very significant specimens from Paleogene localities in central Patagonia (Argentina) attributable to the following taxa: cf. Scaglia kraglievichorum (Barrancan? subage), based on a partial upper molar nearly 60% larger than the type of Scaglia kraglievichorum Simpson, 1957; Astrapotheriidae gen. et sp. 1 (Barrancan? Subage, Middle Eocene), based on an isolated upper molar larger than any other Eocene astrapothere; Astrapotheriidae gen. et sp. 2, based on five isolated upper cheek teeth from “La Cantera” (Gran Barranca, Early Oligocene), characterized by a large, isolated hypocone and accessory cusps on P3-P4. These taxa enlarge the known diversity of Paleogene astrapotheres and document novel evolutionary patterns for these mammals.     

Early Miocene subtropical water temperatures in the southeast Pacific

Cenozoic climate of western South America is strongly controlled by features like Andean uplift and the Humboldt Current. The first strontium isotope age data from central and southern Chile provide a latest Oligocene to late early Miocene age for classic warm-water mollusk faunas reaching as far south as 45°S. Comparison with the biogeography of congeneric living species indicates that sea surface temperatures off central and southern Chile during that time were at least 5 °C higher than today; i.e., minimum annual mean sea surface temperatures for Darwin's Navidad fauna at 34°S are estimated as 20 °C. As expected, the number of tropical taxa decreases towards the south but several are still present as far south as 45°S. The ages scatter relatively broadly between ~ 24 and ~ 16 Ma, partly even within individual localities. Shallow-water and deeper-water faunas are revealed to have similar ages. When considered in light of convincing micropaleontological evidence for late Miocene to early Pliocene depositional ages, the Sr isotope data support a hypothesis that the mollusk fauna is reworked.     

Biogeographic relationships of African carnivoran faunas, 7–1.2 Ma

This study analyses the carnivore component of African fossil faunas from three time slices: 7–5 Ma, 4–3 Ma, and 2.5–1.2 Ma, using cluster analysis and principal coordinate analysis (PCO) of presence/absence data on genera. The faunas mostly cluster by time slice, with the exception of Laetoli (Tanzania) and Ahl al Oughlam (Morocco), which differ from all other faunas. The separation during the Late Miocene of a Chado–Libyan bioprovince from the remainder of Africa is supported. No such distinctions are present in the other time slices. Taxonomic distance is not generally correlated with geographic distance, though if Langebaanweg is removed from the 7–5 Ma time slice, the correlation at that time is significant. Comparison of these paleontological results with phylogeographic studies of modern species leads to some general comments on the analytic power of the fossil record with regard to interregional migrations.     

新疆准噶尔盆地早中新世的最早众古仓鼠及其与稀古仓鼠的对比(英文)

记述了新疆准噶尔盆地早中新世地层中发现的众古仓鼠类化石。材料采集于索索泉组的两个地点,古地磁年龄介于21.9～21.16Ma之间。这是迄今为止中亚和东亚地区众古仓鼠的最早记录。材料可以归入两个种:苏氏众古仓鼠(新种)Democricetodon sui sp.nov.和众古仓鼠未定种Democricetodon?sp.,该未定种材料很少,不足以准确分类。与早中新世其他仓鼠的比较及对稀古仓鼠Spanocricetodon模式种的重新观察表明,苏氏众古仓鼠是中亚地区谢家期的指示性化石,李传夔(1977)所定义的稀古仓鼠为一个有效属,与Democricetodon和Primus属有明显区别;以前归入稀古仓鼠的某些种类应归入其他属。苏氏众古仓鼠与同一地区晚渐新世的Eucricetodon aff.E.caducus形态明显不同,表明它可能不是起源于该地区的古近纪仓鼠,而是迁移至中亚的外来种。     

New Data on Miocene Neotropical Provinciality from Cerdas, Bolivia

We provide the first faunal report for the early/middle Miocene fauna of Cerdas, Bolivia (16.5–15.3 Ma; 20° 52′ S, 66° 19′ W), based primarily on new specimens collected in 2007. As many as twelve species of mammals in nine families are represented. Notoungulates include Palyeidodon obtusum (Toxodontidae), Protypotherium cf. attenuatum and Protypotherium sp. nov. (Interatheriidae), ‘Plesiotypotherium’ minus and possibly Microtypotherium choquecotense (Mesotheriidae), and Hegetotherium? sp. nov. (Hegetotheriidae). Xenarthrans include Stenotatus planus and Prozaedyus sp. (Cingulata: Dasypodidae), Peltephilidae gen. et sp. nov. (Cingulata), and Xyophorus cf. bondesioi (Pilosa: Nothrotheriidae). A new species of litoptern is also present (Macraucheniidae) as well as an unidentified rodent (Chinchillidae: Lagostominae). Two of these Cerdas species occur in Friasian sensu stricto/Colloncuran SALMA faunas of Patagonia, and perhaps one in Santacrucian SALMA faunas. Among middle latitude localities, Cerdas resembles Chucal, Chile (late early Miocene, Santacrucian SALMA) in community composition (e.g., abundant mesotheriids, few rodent species), but has no species in common; it shares one species with Quebrada Honda, Bolivia (middle Miocene, Laventan SALMA), and perhaps as many as three more. These observations indicate that Cerdas is not referable to the Santacrucian, and that the upper limit of this SALMA in the middle latitudes falls somewhere between 17.5 Ma (the top of Chucal) and 16.5 Ma (the base of Cerdas). Based on the range of dates proposed for the youngest Santacrucian intervals in Patagonia, a diachronous offset of up to 2.1 Ma may exist at this point in the SALMA sequence between middle and high latitude faunas.     

A contribution to the chronology of some Old World miocene faunas based on hipparionine horses

The Maragheh Formation, northwestern Iran,provides a local biostratigraphic base upon which to build a biochronology of hipparionine horses that has potential regional importance in ordering faunas of Vallesian and Turolian age in other areas of the Old World. This, as well as faunal and radiometric analysis of the Maragheh sequence, is compared with those aspects of other districts. The radiometric age of the so-called «Hipparion Datum» is 12 Ma, if not slightly older, and more than one species of «Hipparion» may be associated with it. Based on cranial morphology, hipparionine horses of Vallesian age consist of a single group; faunas of Turolian age contain four hipparionine groups by these criteria. A group composed of Hipparion prostylum and later members appears to be the most useful in compiling a possible biochronology. This group may have had an endemic European rather than allochthonous (American) origin. Based on the various assessments discussed here the following temporal sequence of certain faunal localities is proposed (oldest to youngest): Vallesian-Höwenegg (Germany), Bou Hanifia I (N. Africa) and Hostalets de Pierola (Spain). Early Turolian-Kopran, Lower Maragheh (Iran). Medial Turolian-Mont Luberon (France), Kerjabad or Ketschawa, Middle Maragheh (Iran), Saloniki (Macedonia), Pikermi (Greece), Shol'avand, Upper Maragheh (Iran), Samos (Greece). Our assessment of the age of the Mont Luberon and Samos faunas differs from conventional assignments.     

临夏盆地早中新世上庄组的巨獠犀门齿化石(英文)

记述了在临夏盆地早中新世地层中发现的兰州巨獠犀(Aprotodon lanzhouensis)的下门齿化石,其特点为非常粗壮并强烈弯曲。新材料的发现使巨獠犀在临夏盆地的延续时代跨越渐新世/中新世界线的推测得到完全证实。巨獠犀分布的地质时代和地理范围与巨犀重合,但巨獠犀的化石地点和个体数量都相当稀少。巨獠犀的下颌形态功能特点指示其生活于晚始新世至早中新世中国西北、南亚和中亚干旱环境地带中镶嵌分布的少量近水环境。巨獠犀在中中新世之前彻底绝灭,其原因可能是气候变化的结果,也说明临夏盆地早中新世的环境特征与晚渐新世的疏林系统相似,而不同于中中新世的茂密森林。