基于竞争和扩散能力的非捕食-被捕食集合种群系统的竞争机制

对于非捕食 被捕食(食饵)生态系统,强弱物种之间存在一定的竞争影响.在不考虑栖息地毁坏的情况下,引进双向竞争机制,将Tilman的单向竞争模式推广为n集合种群双向竞争模型,并对6-集合种群的竞争动态进行了计算机模拟研究.结果表明,在平衡态,种群竞争共存的条件是其竞争能力与扩散能力呈现指数型负相关关系,竞争的结果使物种的强弱序列发生变化;物种竞争排除与共存受迁移扩散能力和竞争能力影响很大,在局域斑块上竞争排斥的集合种群在广域尺度上可以竞争共存,即逃亡共存.     

昆虫种群空间格局研究方法的最近进展

<正> 昆虫种群空间格局是由物种的行为内禀特征与栖境相互作用而形成的种群个体在空间的分布状况。它的研究无论在生态学理论,还是生产实践上都具有重要意义。空间格局的研究方法可归纳为3类:(1)频次分布拟合x~2检验法;(2)扩散型指数法;(3)回归模型分析法。随着对大量昆虫种群空间格局的实际研究,     

扩散对破碎化景观上宿主-寄生种群动态的影响

景观破碎化和扩散是空间种群模型的重要因素,对生物入侵存在着深远的影响。本章将基于偶对近似模型,探讨由局部和全局宿主-寄生相互作用共同决定的扩散模式对破坏性景观上疾病入侵与传播的影响。其中,生境破坏由生境丧失量与生境破碎化程度来描述。模拟结果显示,宿主和病毒的全局扩散对疾病的入侵与种群密度产生不对称效应：病毒的全局扩散对系统产生的影响较宿主的全局扩散更为显著。不同扩散模式下,生境丧失越高或破碎化程度越低,均将越有害于寄生病毒的入侵;同时,生境的破坏程度也显著地影响了入侵阈值对扩散模式的响应机制。本文研究结果暗示,景观破碎化的空间分布格局以及病毒扩散的限制均可作为物种保护与管理中有效的疾病控制策略。该研究结果在一定意义上丰富和发展了寄生感染理论,为物种保护提供了生态学理论依据。     

祁连山红腹牧草蝗雌雄虫空间格局及关联性

应用地统计学的方法,研究了祁连山北坡石羊河上游,6—10月之间红腹牧草蝗Omocestus haemorrhoidalis(Charpentie)雌雄虫的空间分布格局、动态变化及空间相关性。结果表明:红腹牧草蝗雌雄虫的半变异函数为球状模型与高斯模型,二者空间分布类型相同,均呈聚集分布;随时间的变化均呈现"扩散-聚集-扩散-聚集"的动态特征,7—8月之间雌虫的变化强度明显强于雄虫;二者的模糊贴近度指数在0.53～0.71之间,且指数随红腹牧草蝗的生长呈逐渐增加到减少的变化特征,表明雌雄虫在数量与空间上有较强的跟随关系,且跟随关系随红腹牧草蝗的生长发生明显的阶段性变化。     

羊草与其主要伴生种竞争与共存的格局分析

在羊草种群与其它植物种群的交错区,应用频度、格避形式,格局强度指数对羊草及其主要伴生种之间的共存格局进行了分析。结果表明,羊草及其主要伴生种的格局呈多样化,集聚格局形式是羊草抵御外来物种入侵,或者是自身扩散的一种对策,羊草与其主要伴生种之间存在竞争与共存作用,羊草与芦苇之间通过拮抗作用实现竞争与共存,羊草与鸡儿肠通过竞争而实现共存,光稃茅香,碱茅以营养繁殖策略实现与羊草竞争,指子茅的生长受羊草竞争的抑制。     

应用细胞自动机对一年生植物种群扩散进行理论研究

应用细胞自动机方法构造了用于研究一年生植物扩散的理论模型并应用该模型模拟了一年生植物（杂草）种群在同质环境中的扩散。一年生杂草种子的扩散距离和分布是其种群扩散的主要方式,故本文将其种子扩散分布作为构建邻域细胞函数的基础。根据Howard(1991)^[1]提供的某一年生杂草种子的扩散数据,本文导出了一个25邻的邻域细胞函数和相关的转移函数。建立了一个受控的细胞自动机模型。通过模拟,发现在同质环境中聚集在一起的一年生植物杂草越多就需要越大的控制力才能限制它们扩散;生长于农田边缘的杂草更容易被控制。这些模拟结果表明该模型能较好地表现生态学中的两个众所周知的现象;生物的聚集效应和边缘效应。希望自动机方法和在本文获得的知识将有助于我们制订植物种群的最优管理策略。     

云南滇池入侵虾虎鱼类的共存策略

虾虎鱼类是入侵种的一个重要来源,常数种共同入侵同一区域,其种间如何实现共存是当前鱼类入侵生态学的一个重要科学问题。本研究通过形态分析、空间分布调查及摄食分析等方法,从形态分化和生态位分化角度揭示滇池新入侵的粘皮鯔虾虎鱼与先期入侵近缘种的共存策略。结果表明: 粘皮鯔虾虎鱼与小黄黝鱼、子陵吻虾虎鱼、波氏吻虾虎鱼等先期入侵种在形态上存在显著差异。其空间分布在研究区域上与上述物种存在显著差异,空间生态位指数显著高于其余物种。摄食分析显示,粘皮鯔虾虎鱼的食物多样性指数显著高于其余物种;其主要摄食种类为圆形盘肠溞和腺介虫等,与其他虾虎鱼类主要摄食种类也存在明显差异。在摄食生态位上,滇池入侵虾虎鱼类存在显著分化。这说明形态差异、空间生态位分化及营养生态位分化降低了种间竞争强度,促进了粘皮鯔虾虎鱼与先期入侵虾虎鱼类实现共存。粘皮鯔虾虎鱼可能凭借摄食策略上的优势在种群数量、占据栖息地范围上优于竞争对手,成为研究区域入侵虾虎鱼类中的优势种群。     

不同迁移能力和竞争能力的集合种群竞争模式及其模拟

种群竞争是影响生态系统演化的莺要生态过程之一。本文是在徐彩琳和Tilman研究工作的基础上,将竞争系数引入集合种群动力模式,建立了集合种群之间竞争的数学模型,并对5-集合种群的竞争动态进行了计算机模拟研究。结果表明：种群竞争排除与共存受迁移扩散能力和竞争能力影响很大,排除原理在理论上是存在的,在广域集合种群和实际中物种是竞争共存的．共存的条件是其竞争能力与扩散能力呈非线性负相关关系,竞争的结果使物种的强弱序列发生变化。     

物种竞争的动力机制及数值模拟分析

在徐彩琳和Tilman研究工作的基础上,将竞争系数引入集合种群动力模式,建立了集合种群物种之间竞争的数学模型,并对集合种群5-物种的竞争动态进行了计算机模拟研究．结果表明：物种竞争排除与共存受迁移扩散能力和竞争能力影响很大,排除原理在理论上是存在的,在广域集合种群和实际中物种是竞争共存的,共存的条件是其竞争能力与扩散能力呈非线性负相关关系,竞争的结果使物种的强弱序列发生变化．     

北美车前(Plantago Virginica)和车前(P.asiatica)分布格局的分形分析

北美车前 (ＰｌａｎｔａｇｏｖｒｉｇｉｎｉｃａＬ .)是我国东南地区的一种外来杂草 ,该杂草在浙江、江苏、上海、江西等省区有分布 ,多见于田梗路边、住宅四周、抛荒地、公路两侧等人畜出没频繁的地段 ,其种群生长迅速 ,呈现出生态爆发态势 ,是一种典型的生态入侵种。分形理论已被广泛地应用于自然科学、社会科学中的众多领域。在生态学领域 ,应用分形维数能够客观地表达种群的分布式样 ,同时也能反映种群占有空间、资源利用的能力[1～ 6 ] 。本研究以新入侵种北美车前为对象 ,并与我国境内分布较广的杂草车前 (Ｐ .ａｓｉａｔｉｃａ)作比…     

Uneven distribution of weeds along extensive transects in Australia’s Northern Territory points to management solutions

Summary The Top End region of the Northern Territory, Australia, is noted for its relatively unmodified natural state. To gain some insight into the potential for maintaining ecosystem health in this region we undertook a study that assessed the distribution of weeds across very extensive transects. This weed survey was distinct from other studies in that many of the sample sites were distant from tracks or other infrastructure. Twenty‐one weed species were recorded along 2000 km of transects. Weeds were reported from 18.7% of the 718 sample points. The incidence of weeds was found to be significantly associated with land tenure, being highest on pastoral lands and peri‐urban areas, and very low on Aboriginal lands. The incidence of weeds increased significantly with increasing levels of infrastructure and with increasing proximity to watercourses. There are three main conclusions from this study. First, much of the Top End, particularly remote Aboriginal lands, has exceptionally low levels of weed infestation. Secondly, in such areas, given the relatively small extent of vegetation change through weed invasion, maintenance or re‐imposition of traditional fire regimes should be achievable. Thirdly, there is substantial potential for spread of weeds to remote areas, with such spread most likely to occur through increased penetration by infrastructure. Importantly this study indicates that there is still opportunity to prevent widespread weed invasion across the Top End, which is timely given the current Government consideration of the potential for the region to support future agricultural expansion and the fast‐paced development of mining, oil and gas resources.     

The impact of exotic weed competition on a rare New Zealand outcrop herb,

Nearly one quarter of New Zealand’s unique vascular plant flora is threatened, and weed invasion is implicated in the decline of more than half of these threatened species. However, there is little experimental evidence showing that invasive weeds have a direct impact on threatened native plants. This study experimentally tested the hypothesis that competition with invasive weeds threatens the rare outcrop plant Pachycladon cheesemanii (Brassicaceae). Pachycladon cheesemanii is a threatened South Island, New Zealand endemic with a distribution nearly confined to rock outcrops. It has disappeared from historical record sites throughout its range. The effects of weed competition and habitat on P. cheesemanii establishment, growth and survival were investigated by sowing seed into replicated plots subject to three treatments: weed removal, soil disturbed and unweeded control, in three habitat types: forested and open rock outcrops and open tussock grassland. The experiments were carried out at three locations: Mt Somers (Canterbury), Wye Creek and Diamond Lake (Otago). Within weedy rock outcrop habitat, weed removal significantly increased the rate of P. cheesemanii germination, and appeared to increase seedling growth rates, implying that weeds can negatively impact populations. Relative to rock outcrop habitat, P. cheesemanii germination was very low in adjacent open grassland habitat regardless of weeding treatment. Demographic monitoring of four natural populations of P. cheesemanii revealed that seed production is highly variable among populations and may be limited by browse and mechanical damage to inflorescences. Pachycladon cheesemanii does produce a persistent seed bank but most seed is found close to parent populations. Our results suggest that competition with invading weeds threatens current P. cheesemanii populations, that plant establishment can be enhanced by weed removal, and that considerable potential exists for artificially expanding populations by sowing seed into appropriate weed-free habitat.     

Spatial interplay of plant competition and consumer foraging mediate plant coexistence and drive the invasion ratchet

Indirect effects may play an important role in structuring plant communities. Using a spatially explicit model of consumer foraging and plant competition, we demonstrate how the relationship between the spatial area over which plants compete and the spatial scale of consumer behaviour can determine the outcome of competition when one plant species provides a refuge for mobile consumers (i.e. refuge-mediated apparent competition). Once an initial population of the invader is established, complete invasion may be inevitable because of an ever-advancing invasion front ratchets forward driven by a feeding front of mobile consumers. Because the spatial extent of apparent competition determines the area available for colonization, consumers may also dictate the rate at which an invasion occurs. We find that, as long as refuge-mediated apparent competition is sufficiently localized, invasion is possible even in systems characterized by low overall levels of consumer pressure. Moreover, we show that a stable equilibrium can result in which both resident and invading plants coexist, suggesting that spatial heterogeneity created by refuge-mediated apparent competition may be important in mediating coexistence in plant communities. The spatial interplay of consumer behaviour and plant competition may be an underappreciated mechanism affecting the composition, diversity and spatial pattern of plant communities.     

Analysing spatial correlation of weeds and harvester ants in cereal fields using point processes

The interaction between the spatial distribution of weed richness and weed cover and the spatial location of harvester ant nets was investigated in cereal fields. The understanding of such interdependencies can be relevant to understand weed population dynamics in dryland cereal fields and may enhance management strategies for weed control. We used spatial statistical tools derived from point process theory. In particular, we compared the two spatial configurations by assuming two different point patterns. We did so by replacing the random weed fields by a related point pattern and comparing it with the point pattern of harvester ants. Our results suggest that areas with a high density of ant nests are, in this case study, in areas with low weed richness and that large nests have a greater impact than small nests. Considering that only one field was analysed, preserving and enhancing regular ant nest distributions, especially of large nests, might have an impact on depleting weeds and consequently enhancing weed control.     

Does weed suppression by high crop density depend on crop spatial pattern and soil water availability?

Size-asymmetric competition, in which larger plants obtain a disproportionally larger share of contested resources, can be applied in agriculture to suppress weeds by increasing crop density and spatial uniformity, as these practices enhance the initial size-asymmetric competitive advantages of crop seedlings over weed seedlings early in the growing season. We do not yet know how agronomic factors influence weed suppression at high crop density. We performed a field experiment to ask how crop density, spatial pattern and irrigation interact to influence weed suppression and grain yield in semi-arid croplands. The experimental was a factorial design with 4 factors: wheat cultivar (Ningchun4, Xihan2), irrigation level (control, irrigated), sowing density (low, 196 seeds m⁻²; moderate, 400 seeds m^-2; high, 625 seeds m⁻²), and spatial sowing pattern (rows, uniform). Weed growth was effectively suppressed by increased crop density and spatial uniformity. Effects of crop density on weed suppression and grain yield were more pronounced in the uniform pattern than in crop rows. Weed biomass was 55.7% lower and grain yield increased 29.7% higher in the high density uniform pattern compared to the low density and row pattern. Crop density interacted with cultivar in determining both weed biomass and grain yield, potentially reflecting different traits regulating crop competitive ability. Irrigation and crop density had additive effects on weed biomass but interacted to influence grain yield. Our findings support the idea that increased crop density and spatial uniformity can make a valuable and environmentally friendly contribution to weed control in wheat, reducing the need for chemical or mechanical weed control. We need a better understanding of the interactions among climate, agricultural management and crop genotype to improve our ability to effectively suppress weeds with high crop density and spatial uniformity.     

Controlling infestations of Parkinsonia aculeata in a riparian zone at the landscape scale

Abstract To prioritize weed management activities it is necessary to predict the extent of future infestations of individual weed species. Management effort should focus on populations which are likely to spread and produce new satellite populations. Deciding which aspects of an invading population to manage, new emerging distant satellites or the existing core from which most seed originates has been the subject of much debate. We use extensive survey data to define the core and satellite populations and dispersal function of Parkinsonia aculeata in the semi‐arid rangelands of the Northern Territory of Australia. We described populations in terms of three size classes, classified on the basis of plant height (small (<0.4 m), medium (>0.4 and <2.5 m) and large (>2.5 m)). Data were recorded as a spatial point pattern and a dispersal function was derived from the distribution of the distance of each medium plant to the nearest P. aculeata neighbour that was greater than 2.5 m. All large and medium P. aculeata were classified as members of core or satellite populations. From these classifications, we tested critical assumptions of Moody and Mack's model, namely (i) the satellites form multiple disjunct foci that do not overlap; (ii) the system is homogenous and there are no abiotic or biotic restrictions to the weeds expansion enabling it to expand radially in any direction; (iii) the densities of all foci are uniform; (iv) there is no extinction through stochastic processes; and (v) there are no threshold effects. When the medium size class was considered, the population density and area occupied by the weed increased. Two satellite populations were comprised solely of medium plants, suggesting the number of satellites increased marginally from eight to 10 when data from both size classes were considered. Most individuals were part of the core population. The core population increased in density and we argue this core population is most likely to give rise to new satellites and should be managed in preference to emerging satellite populations. We suggest the invasion is behaving like a source‐sink system, where the core population is the source of most new medium plants.     

Competition between similar invasive species: modeling invasional interference across a landscape

As the number of biological invasions increases, interactions between different invasive species will become increasingly important. Several studies have examined facilitative invader–invader interactions, potentially leading to invasional meltdown. However, if invader interactions are negative, invasional interference may lead to lower invader abundance and spread. To explore this possibility, we develop models of two competing invaders. A landscape simulation model examines the patterns created by two such species invading into the same region. We then apply the model to a case study of Carduus nutans L. and C. acanthoides L., two economically important invasive weeds that exhibit a spatially segregated distribution in central Pennsylvania, USA. The results of these spatially-explicit models are generally consistent with the results of classic Lotka–Volterra competition models, with widespread coexistence predicted if interspecific effects are weaker than intraspecific effects for both species. However, spatial segregation of the two species (with lower net densities and no further spread) may arise, particularly when interspecific competition is stronger than intraspecific competition. A moving area of overlap may result when one species is a superior competitor. In the Carduus system, our model suggests that invasional interference will lead to lower levels of each species when together, but a similar net level of thistle invasion due to the similarity of intra- and interspecific competition. Thus, invasional interference may have important implications for the distribution and management of invasive species.     

Invasion and adaptive evolution for individual-based spatially structured populations

The interplay between space and evolution is an important issue in population dynamics, that is particularly crucial in the emergence of polymorphism and spatial patterns. Recently, biological studies suggest that invasion and evolution are closely related. Here, we model the interplay between space and evolution starting with an individual-based approach and show the important role of parameter scalings on clustering and invasion. We consider a stochastic discrete model with birth, death, competition, mutation and spatial diffusion, where all the parameters may depend both on the position and on the phenotypic trait of individuals. The spatial motion is driven by a reflected diffusion in a bounded domain. The interaction is modelled as a trait competition between individuals within a given spatial interaction range. First, we give an algorithmic construction of the process. Next, we obtain large population approximations, as weak solutions of nonlinear reaction–diffusion equations. As the spatial interaction range is fixed, the nonlinearity is nonlocal. Then, we make the interaction range decrease to zero and prove the convergence to spatially localized nonlinear reaction–diffusion equations. Finally, a discussion of three concrete examples is proposed, based on simulations of the microscopic individual-based model. These examples illustrate the strong effects of the spatial interaction range on the emergence of spatial and phenotypic diversity (clustering and polymorphism) and on the interplay between invasion and evolution. The simulations focus on the qualitative differences between local and nonlocal interactions.      

沟叶结缕草草坪杂草群落的时空动态

杂草群落结构的异质性是群落动态的主要属性.基于物种频度和幂函数法则,对沟叶结缕草草坪杂草群落在时间维度上的物种结构和空间异质性的变化进行了研究.结果表明： 从2007到2009年,沟叶结缕草草坪3年共发生杂草43种,隶属19科,其中,禾本科、菊科、莎草科和茜草科杂草在草坪杂草中占据优势,多年生杂草占杂草比例最大,并呈逐年上升趋势;杂草群落具有比随机分布更高的空间异质性;杂草优势种年内随季节变化明显,表现出“双子叶杂草－单子叶杂草－双子叶杂草”、“多年生杂草－一年生杂草－多年生杂草”交替出现的规律;杂草群落的空间异质性在夏季大于冬季,多样性和均匀性在夏秋季大于冬春季,在夏季表现出高异质性的杂草种类比其他季节多;杂草群落空间异质性、多样性年际间变化不显著,均匀性呈逐年降低趋势.
     

Early establishment of planted Retama sphaerocarpa seedlings under different levels of light,water and weed competition

Large amounts of former cropland are being abandoned in developedregions. To formulate guidelines for land reclamation programmes, we exploredthe effects of artificial shading, irrigation, and removal of weed competitionon the performance of Retama sphaerocarpa (L.) Boiss.seedlings in a factorial experiment located in an abandoned cropland in CentralSpain. R. sphaerocarpa is of interest for revegetationbecause it is a drought tolerant leguminous shrub that is a major structuralcomponent of the native plant community. Seedling performance was evaluated inthree ways: seedling survivorship, growth, and photochemical efficiency. Wealsomeasured soil moisture and weed biomass production and found that bothincreasedunder artificial shading conditions. Soil moisture increased very slightlywhereweeds were removed. Thus, increased transpiration from weeds outweighed reducedevaporation from soils due to shading by weeds. Artificial shading was the mosteffective treatment for seedling survivorship, followed by removal ofcompetition by weeds. After summer, 34 % of the seedlings survived in the mostfavourable conditions (artificially shaded plots where weeds were removed),compared to ca. 1 % in full-light plots with no removal of weed competition. Apositive effect of irrigation was found for growth of seedling cover and heightin shaded plots. The analysis of photochemical efficiency pointed out therelevance of weed competition removal, and confirmed the usefulness of fastfluorescence transient techniques for the quantification of seedlingperformance. The data suggest that competition between seedlings and weeds wasprimarily for water rather than for light. We conclude that i) artificialshading improved seedling performance, but this is a little practical techniquebecause of its cost; ii) as weeds compete with, rather than facilitate, plantedseedlings, weed clipping around the seedlings is a feasible technique thatwouldimprove seedling survival; and iii) seedling performance could alsoconsiderablyimprove with a higher irrigation than was used in this experiment(75lm^–2 per growth period), provided that weedsare removed.