New Harpacticoida (Crustacea, Copepoda) from the North Atlantic Ocean. V. Three New Species of Metahuntemannia Smirnov (Cletodidae)

Metahuntemannia triarticulata sp.n., M. arctica sp.n. and M. bifida sp.n. arc described. M. triurficulata and M. arctica belong to the 'spinosa' group within the genus Metahuntemannia Smirnov. From the closely related species, M. triarticulata differs in the possession of 3-segmented Enp P3, M. arctica in the segmentation of the Al, and both new species in the setation of their swimming legs P1–P5. M. hifida belongs to the 'talpa' group within Metahuntemannia and differs from the closely related species in the setation of the swimming legs P2–P4. The species relationships and the phylogeny within the genus are discussed. All species were collected at the Iceland-Faroe Ridge from depths between 435 and 2500 m.     

Composition of the genera <Emphasis Type="Italic">Gnaptorina</Emphasis> Reitter and <Emphasis Type="Italic">Pseudognaptorina</Emphasis> Kaszab of the tribe Blaptini (Coleoptera,Tenebrionidae)

Based on the structure of the segments of the male fore and middle tarsi, three subgenera are distinguished in the genus Gnaptorina Reitter: Gnaptorina s. str. (brushes or tufts of pale setae are present on the sole surface of the 1st–3rd segments of the fore tarsus and also on the 1st and 2nd segments of the middle tarsus, the type species Gnaptorina felicitana Reitter, 1887), Boreoptorina subgen. n. (a brush is present on the sole surface of the 1st segment of the fore tarsus, but is absent on the sole surface of the segments of middle tarsus, the inner margin of the male fore tibia with a hairy brush, the type species Gnaptorina cordicollis G. Medvedev, 1998), and Hesperoptorina subgen. n. (a flat hairy brush is present only on the sole surface of the 1st segment of the fore tarsus, type species Gnaptorina brucei Blair, 1923). The genus Pseudognaptorina Kaszab, 1977 is considered monotypical. Lectotypes of four species of Gnaptorina are designated.     

Phylogeny and zoogeography of three new species of the genus Hexabathynella (Crustacea, Malacostraca, Bathynellacea) from North America

Three new species of the genus Hexabathynella , H. otayana sp. n., H. hessleri sp. n. and H. muliebris sp. n., from North America are described. A comparison with previous studies results in the reevaluation of characters already known and the discovery of new constitutive characters. The distal spine of the mandibular spine row is regarded as a synapomorphy of the three new species and two previously known species, H. knoepffleri , H. minuta , which constitute a monophyletic group within the genus, the knoepffleri-group. The phylogenetic relationships between the five species are depicted with a cladogram. The setation on the uropodal exopod is shown to be not only a new unique character for the genus, but also a significant one in classifying its species. The presence of the ancestor species of the genus before the Triassic and dispersal during Mesozoic continental drift are assumed to explain the transatlantic occurrence of the knoepffleri-group and the distribution pattern of the other species.     

Two New species of the dipteran genus Dolichopus Latreille (Diptera,Dolichopodidae) from Siberia and Mongolia

Two new species of the genus Dolichopus Latreille (Diptera, Dolichopodidae) are described from Mongolia and Siberia. Dolichopus tschernovi Negrobov, Barkalov et Selivanova, sp. n. is similar to Dolichopus bayaticus Negrobov, 1976, but differs in the presence of one seta on the hind metatarsus, the shorter 3rd antennal segment, and in the structure of the hypopygium. Dolichopus lenensis Negrobov, Barkalov et Selivanova, sp. n. resembles Dolichopus setitarsis Negrobov et Barkalov, 1977, but differs in the slightly widened 2nd–4th segments of the middle tarsus, in the presence of one seta on the hind metatarsus, and in the structure of the hypopygium. Drawings of the antennae, tarsi, and hypopygium of the new species are given. The holotypes of the new species are deposited in the Zoological Institute, the Russian Academy of Sciences (St. Petersburg).     

Revision of genus Melamphaes (Melamphaidae): Part 4. Multi-raker species: M. ebelingi, M. occlusus sp. nova, and M. nikolayi sp. nova

In the fourth part of the revision of the genus Melamphaes (Melamphaidae) multi-raker species (20 and more rakers at the first gill arch, rarely 19??in M. ebelingi) with eight soft rays in the ventral fin are considered. In this species, temporal spine is absent, there are 15 rays in the pectoral fin, and there are 11 to 12 abdominal vertebrae. M. ebelingi inhabits tropical and subtropical waters of the Atlantic Ocean. A new catch of this rare species in the northeastern part of the ocean is reported. The new species M. occlusus is described from one specimen from the subtropical part of the southwestern part of the Indian Ocean. Still another new species, M. nikolayi, is described from one specimen from the northwestern part of the Pacific Ocean (subtropical waters). A table for identification of 15 multi-raker species of the genus Melamphaes is provided.     

Four new Mouse Spider species (Araneae,Mygalomorphae, Actinopodidae,Missulena) from Western Australia

Four new species of the Mouse Spider genus Missulena Walckenaer, 1805 (family Actinopodidae) are described from Western Australia based on morphological features of adult males. Missulena leniae sp. n.(from the Carnarvon and Yalgoo biogeographic regions), Missulena mainae sp. n. (Carnarvon), Missulena melissae sp. n. (Pilbara) and Missulena pinguipes sp. n. (Mallee) represent a broad spectrum of morphological diversity found in this genus and differ from other congeners by details of the male copulatory bulb, colour patterns, eye sizes, leg morphology and leg spination. Two of the species, M. pinguipes sp. n. and M. mainae sp. n., are characterised by swollen metatarsi of the fourth legs in males, a feature not previously recorded in the family. A key to males of all named Missulena species from Australia is presented and allows their identification based on external morphology.     

Two new species of Mediomastus (Annelida,Capitellidae) from Tokyo Bay,Japan

Two undescribed species of polychaetes in Mediomastus (Annelida: Capitellidae) were collected from intertidal to shallow habitats in Tokyo Bay, Japan. These are M. duobalteus sp. n. and M. hanedaensis sp. n. Mediomastus duobalteus sp. n. is distinguishable from all congeners by the following characters: 1) segments 3, 4, 8–11 stainable with methyl green, 2) thoracic capillary chaetae unilimbate, 3) abdominal capillary chaetae absent, 4) paddle-like chaetae in the thorax absent, and 5) abdominal hooded hooks not flared. Mediomastus hanedaensis sp. n. is similar to M. warrenae Green, 2002, but differs from the latter in the shapes of the thoracic capillary chaetae and the abdominal hooded hooks, and the staining pattern with methyl green. In addition, a key to all Mediomastus species is provided.     

Psammic Procyclopina (Copepoda, Cyclopoida) from the coast of Brazil

Procyclopina polyarthra Herbst is redescribed and three new species, P. maricopeba, P. feiticeira and P. uguaipuku , are described from intertidal interstitial waters. Procyclopina is distinguished within the family by the enditic setation of the maxilliped 0,1,3,4, the 19-segmented antennule, and three elements in the leg 5 exopod of the female. Procyclopina has its distribution restricted to the coast of Säo Paulo, Brazil. Procyclopina maricopeba sp. n. is distinguished from P. polyarthra by the three-segmented leg 5 in the male and by the length: width ratio of the genital double somite in the female. Procyclopina feiticeira sp. n. is unique in the genus by lacking the inner seta on legs 14 exp-I, and by the incomplete fusion of genital (seventh thoracic) and first abdominal somites in the female. Procyclopina uguaipuku sp. n. is characterized by the antennulary armature, by the caudal ramus (much longer than wide), and by the position of the proximal seta of the leg 4 enp-2 which is set on a protuberance on the posterior face. Morphological characters of Procyclopina are compared with those of other cyclopinids.     

Meristoderes gen. nov., a new kinorhynch genus, with the description of two new species and their implications for echinoderid phylogeny (Kinorhyncha: Cyclorhagida, Echinoderidae)

A new kinorhynch genus, Meristoderes gen. nov., and two new species from Spain and the Solomon Islands, respectively, are described. The new genus is distinguished from all other genera by the first segment consisting of a closed cuticular ring, and the second segment having partial tergosternal junctions, and a superficial midventral fold. This is a new cuticular configuration that may shed light into the phylogenetic relationships of echinoderid kinorhynchs. Meristoderes macracanthus gen. et sp. nov. from the Mediterranean coast of Spain is recognised by the presence of middorsal spines on segments 4, 6 and 8, ventrolateral tubules on segment 2, lateroventral tubules on segment 5, lateroventral spines on segments 6-9, lateral accessory tubules on segment 8, one pair of laterodorsal tubules on segment 10. Meristoderes galatheae sp. nov. from the Solomon Islands is recognized by having a middorsal spine on segment 4 only, ventrolateral tubules on segment 2, lateroventral tubules on segment 5, lateroventral spines on segments 6-9, lateral accessory tubules on segment 8 and subdorsal tubules on segment 10. Both species have a pattern of paraventral perforation site clusters on segments 3-9, with conspicuously long bracteate hairs from the posteriormost perforations sites on the segments 3-7 and 3-6, respectively.The new genus Meristoderes gen. nov. is included into the family Echinoderidae Bütschli, 1876 and appears closely related with the genera Cephalorhyncha Adrianov, 1999 and Echinoderes Claparède, 1863. The new information it provides is discussed to clarify the internal phylogeny of Echinoderidae. The terminology for cuticular characters in the overlapping area between consecutive segments is also standardized.     

Paratullbergia Womersley in China: the description of a new species and a key to the genus (Collembola,Tullbergiidae)

The genus Paratullbergia Womersley, 1930 is recorded for the first time from China. Paratullbergia changfengensis sp. n. from Shanghai is described and illustrated. It is characterized by the presence of 1+1 pseudocelli on thoracic segment I, with two pairs of pseudocelli on each of thoracic segments II and III, presence of seta px on abdominal segment IV, seta a2 and p4 on abdominal segment V as microsetae, and less differentiated sensory seta p3 on abdominal segment V. Both sexes present. The new species can be easily distinguished from its congeners by the presence of pseudocelli on thoracic segment I. An updated key to the world species of the genus Paratullbergia is provided.     

Description of Eucyclops tziscao sp. n., E. angeli sp. n., and a new record of E. festivus Lindberg, 1955 (Cyclopoida,Cyclopidae, Eucyclopinae) in Chiapas,Mexico

Two new species of the freshwater cyclopoid genera Eucyclops are described, Eucyclops tziscao sp. n. and E. angeli sp. n. Both species belong to the serrulatus-group defined by morphological features such as: the presence of distal spinules or hair-like setae (groups N1 and N2) on frontal surface of antennal basis; the fourth leg coxa with a strong inner spine that bears dense setules on inner side, yet proximally naked (large gap) on outer side; and a 12-segmented antennule with smooth hyaline membrane on the three distalmost segments. Eucyclops tziscao sp. n. is morphologically similar to E. bondi and E. conrowae but differs from these species in having a unique combination of characters, including a caudal ramus 4.05±0.25 times as long as wide, lateral seta of Enp3P4 modified as a strong, sclerotized blunt seta, coxal spine of fourth leg with inner spinule-like setules distally, and sixth leg of males bearing a strong and long inner spine 2.3 times longer than median seta. Eucyclops angeli sp. n. can be distinguished by an unique combination of morphological features: the short caudal ramus; the long spine on the sixth antennular segment of A1; the presence of one additional group of spinules (N12’) on the caudal surface of A2; the presence of long setae in females, or short spinules in males on the lateral margin of fourth prosomite; the strong ornamentation of the intercoxal sclerite of P4, specially group I modified as long denticles; the distal modified setae of Exp3P3 and Exp3P4 in females and males; and the short lateral seta of P5. Finally, we report on a new record of E. festivus in México, and add data on morphology of the species.     

A new species of Odaginiceps Fiers, 1995 (Copepoda, Harpacticoida, Tetragonicipitidae) from the Mediterranean coast of Turkey

Male and female of Odaginiceps korykosensis sp. n. (Copepoda, Harpacticoida, Tetragonicipitidae), collected in the intertidal zone of Kızkalesi beach along the Mediterranean coast of Turkey (Mersin Province), are described. The new species is the fifth member of the genus and can easily be distinguished from the other species by the presence of four setae/spines on the second endopodal segment of P4 and by the structure of the caudal rami. Previously, representatives of the genus Odaginiceps have been reported from Gulf of Mexico, off Bermuda and Kenya. Odaginiceps korykosensis sp. n. is the first record of the genus in the Mediterranean Sea.     

Revision of the genusAcanthaphritis (Percophidae) with the description of a new species

The percophid genusAcanthaphritis is reviewed.Spinapsaron andBranchiopsaron are synonymized withAcanthaphritis. The genus is characterized by a barbel on the snout tip in males, a single maxillary spine, cheek scales, ctenoid lateral line scales, prevomerine teeth present, palatine teeth absent, posterodorsal edge of opercular membrane ragged, five dorsal spines and seven branchiostegal rays. Four species (one new) are recognised in the genus:A. grandisquamis, A. barbata, A. ozawai andA. unoorum n. sp. A lectotype is designated forA. grandisquamis. The new species,A. unoorum, is characterized by having 23–25 dorsal fin rays, 28–30 anal fin rays, 37–39 lateral line scales, 37–39 vertebrae, one or two brown spots on the opercle, two longitudinal rows of yellow marks on the body when fresh, two longitudinal dark lines and a longitudinal row of dark marks on the body in males, and a single longitudinal dark line and longitudinal row of dark marks on the body in females.     

On the fauna of bristletails of the genera <Emphasis Type="Italic">Petrobius</Emphasis> and <Emphasis Type="Italic">Trigoniophthalmus</Emphasis> (Thysanura,Machilidae) from the Caucasus

Five new species, Petrobius caucasicus sp. n., Trigoniophthalmus presimplex sp. n., T. divnogorski sp. n., T. kislovodski sp. n., and T. adigei sp. n. (Machilidae), are described from the Caucasus. A key to all the known species of the genus Trigoniophthalmus is given. Two subgenera (Trigoniophthalmus s. str., Trigoniocellus subgen. n.) in the genus Trigoniophthalmus are described. 2 + 2 retractile vesicles are present on abdominal segments II–IV in the species of Trigoniocellus subgen. n. and on II–V abdominal segments in Trigoniophthalmus s. str. Analysis of the evolution and phylogenetic relations between the species of the genus Trigoniophthalmus is performed for the first time. The formation of bristletails of the genus Trigoniophthalmus appears to occur in the Caucasian mountain province of the European area, from where they penetrated to the Central European mixed province and the Mediterranean subarea. Among the described species of the genus, T. kislovodski is distinguished by the greatest number of plesiomorphic states of morphological characters.     

Apodopsyllus gabesensis n. sp.: a new species of Paramesochridae (Copepoda: Harpacticoida) from the Gulf of Gabès (south-eastern Tunisia)

Several specimens belonging to a new species of Apodopsyllus were collected during a study on the diversity of meiobenthic communities in the Gulf of Gabès, a Mediterranean shallow-water bay at the south-eastern coast of Tunisia in July 2005. The new species Apodopsyllus gabesensis n. sp. shares the characteristics of the genus such as the lack of endopods from P2 to P4 and the soft and slightly cuticularized body. Apodopsyllus gabesensis n. sp. belongs to the few known species of Apodopsyllus that are described to have comparably distinct patterns of dorsal and lateral cuticular plates and pores. Besides a typical combination of characters that clearly distinguishes the new species from its congeners, the new species shows the following unique single characters: female P5 with a hitherto unknown combination of shape of the exopodal part and shape and setation of the baseoendopodal lobe with two small stout spines; male P6 with a particular shape, and a distinct armature of the exopodal spines in P2–P4 in the male that are pinnate with very short spinules contrary to the female where spines are smooth. The genus Apodopsyllus contains 26 species with the inclusion of the new species.     

A new parasitic copepod (Cyclopoida: Bomolochidae) from a ponyfish (Leiognathidae) caught in Egyptian Mediterranean waters,with a review of hosts and key to species of Nothobomolochus

A new bomolochid copepod belonging to the genus Nothobomolochus Vervoort, 1962 is described from a Red Sea fish species, a ponyfish of the family Leiognathidae that has become established in the Eastern Mediterranean. The new species, N. leiognathicola n. sp., is based on material obtained from the gill chamber of the Red Sea immigrant ponyfish Leiognathus klunzingeri (Steindachner), caught in Egyptian waters off the Alexandria coast at Abuqir. A second new species, N. monodi n. sp., is established to accommodate some material previously described as N. denticulatus (Bassett-Smith, 1898), from the host Hemiramphus far Forsskål. A review of host records reveals that Nothobomolochus species utilise hosts representing five different orders, but are most commonly found on beloniform, clupeiform and perciform fishes. A newly constructed key to the 37 valid species of Nothobomolochus is presented.     

Filaires Splendidofilariinae de lézards: nouvelles espèces,redescription, cycle chez phlébotome

Four new species of Splendidofilariinae from different families of lizards are described: Thamugadia wertheimae n. sp., from Acanthodactylus b. boskianus from Israel, Madathamugadia versterae n. sp., from Mabuya capensis from the Northern Transvaal; M. bissani n. sp., from Mabuya quinquetaeniata from Mali; and M. ineichi n. sp., from Pseudocordylus microlepidotus melanotus from the Orange Free State. The life-cycle of M. ineichi is described. It occurs in a Phlebotomus species and is similar to the two known life-cycles in the genus. The new combination Madathamugadia huambensis (Petit, Bain, Gomes & Touratier, 1983) is proposed for the filarial worm from Mabuya aff. quinquetaeniata in Angola, previously placed in the genus Piratuboides (Oswaldofilariinae). Pseudothamugadia physignathi, from Physignathus lesueurii in Australia, is redescribed. In New Guinea, this filarid, or a closely related species, is a parasite of Chlamydosaurus kingii. The presence of splendidofilariines in lizards from southern and western Africa is a new finding. These Ethiopian species show several original characteristics but exhibit affinities with the two Madathamugadia spp. from Malagasy. M. ineichi appears to be a primitive form with a well-developed buccal capsule and well-developed pre- and post-cloacal papillae. Two lines appear to derive from this group: the first is characterised by the reduction of the post-cloacal papillae and comprises the eight species of Madathamugadia in the Ethiopian and Mediterranean regions; the second line is characterised by a reduction of the pre-cloacal papillae and includes the four species of Thamugadia in the Mediterranean region. Morphologically, Pseudothamugadia appears to be closely related to Madathamugadia.     

An unusual new species of paguroid (Crustacea,Anomura, Paguridae) from deep waters of the Gulf of Mexico

A new hermit crab species of the family Paguridae, Tomopaguropsis ahkinpechensis sp. n., is described from deep waters (780–827 m) of the Gulf of Mexico. This is the second species of Tomopaguropsis known from the western Atlantic, and the fifth worldwide. The new species is morphologically most similar to a species from Indonesia, Tomopaguropsis crinita McLaughlin, 1997, the two having ocular peduncles that diminish in width distally, reduced corneas, dense cheliped setation, and males lacking paired pleopods 1. The calcified figs on the branchiostegite and anterodorsally on the posterior carapace, and the calcified first pleonal somite that is not fused to the last thoracic somite, are unusual paguroid characters. A discussion of the affinities and characters that define this new species is included, along with a key to all five species of Tomopaguropsis.