浙江桐庐早二叠世船山组介形类化石

研究报道的介形类化石系采自浙江省桐庐县瑶林镇沈村剖面早二叠世船山组,计14 属4 亚属22 种,包括4新种和4未定种。其中12属4亚属和19种产自船山组第三段,并被归为Hollinella(Praehollinella) ema ciata Basslerella ola 组合,代表早二叠世船山世隆林期。牙形刺化石主要产自船山组第一段,带化石Streptogna thodus elegantulus 及其共生的其它牙形刺化石,指明船山组第一段的时代为中、晚卡西莫夫期到早格泽里期。根据介形类动物群生态组合和岩石特征,推测船山组第三段是在温暖的近岸浅水且为低能的开阔海环境下沉积的。     

华南法门期牙形刺的辐射

F／F牙形刺灭绝事件以后,牙形刺很快进入复苏期（王成源、Ziegler,2004）。有关法门期牙形刺的辐射,在国内外的文献中还没有详细的论述或论证。本文基于以前对广西桂林垌村和龙门剖面的牙形刺集群灭绝与复苏的研究,2003年又在垌村剖面加密采样,重点是研究F／F事件后,牙形刺由复苏到辐射阶段的特征,探索牙形刺何时和如何由复苏期进化到辐射期。本文的材料主要是依据华南的资料,特别是本文作者研究多年的垌村剖面的资料。牙形刺的灭绝与复苏是以法门期最主要的属Palmatolepis为主线,在种一级的水平上确定的。法门期牙形刺的辐射同样是以Palmatolepis属内种一级的水平上讨论的。同时也涉及到其它属。本文首次明确提出：牙形刺在晚triangularis带就进入辐射期,因为正是从晚triangularis带,Palmatolepis的minuta,werneri,sandbergi,perlobata,cf．regularis,subperlobata,tenupunctata7个重要种群开始出现,并在法门期中期达到辐射期的高峰。     

滇西保山地区丁家寨组、卧牛寺组牙形刺的时代

本首次描述了滇西保山卧牛寺组及永德丁家寨组的Rabeignathus牙形刺动物群,其中2个新种：Rabeignathus yunnanensis sp．nov．,R．ritterianus sp．nov．,并划分出3个牙形刺带,进一步确定了古生物地层工作争论已久的卧牛寺组的时代为Artinskian晚期-Kungurian早期;丁家寨组上段为Sakmarian晚期-Artinskian早期。该牙形刺动物群为暖温型动物群,结合保山地块当时的古地磁资料,丁家寨组、卧牛寺组沉积之时,应处于边缘冈瓦纳区。卧牛寺组玄武岩喷发时间的确定,预示了保山地块从边缘冈瓦纳区分离出来的时间为Artinskian晚期。     

日本志留纪至三叠纪牙形刺生物地层

本文系统地回顾了日本志留纪至三叠纪牙形刺研究的历史和现在的成果。志留纪牙形刺只有少数零星的报道,没有建立化石带;早泥盆世已建立了５个牙形刺组合,没有中、晚泥盆世的记录;石炭纪有８个牙形刺带,其中晚石炭世３个牙形刺带;二叠纪５个牙形刺带或动物群,其中,中、晚二叠世各１个带（动物群）;三叠纪可划分出１４个牙形刺带。     

依据牙形刺确定的内蒙古苏尼特左旗泥盆系泥鳅河组的时代

内蒙古苏尼特左旗泥鳅河组一段下部产Caudicriodus woschmidti hesperius等牙形刺,时代为Lochkovian(洛赫考夫期)期最早期。泥鳅河组二段最下部产牙形刺Masaraella pandora W new morph,Pandorinellinacf.miae等,时代为Pragian(布拉格期)期最早期,或由Lochkovian晚期pandora beta-gilberti带到Pragian期早期的steinachensis带。依据牙形刺确定的泥鳅河组一段—二段的时代为早泥盆世早、中期,即Lochkovian期到Pragian期的沉积。本文还首次确认了Masaraella属在中国的存在,并描述了Masaraella pandora W新的形态型。     

湖北恩施志留系秀山组顶部的牙形刺及其地层意义

本文描述了湖北恩施太阳河灯草坪剖面志留系秀山组顶部灰岩夹层中的牙形刺。秀山组顶部的灰岩产出较丰富的牙形刺化石,共10属14种,包括Pterospathodus eopennatus,P.a.cf.angulatus,Apsidognathus sp.,Ozakodina paraplanussima,Ozakodina sp.,?Distomodus sp.,?Tuberocostadontus sp.,Coryssagnathus shaannanensis,Wurmiella recave,Wurmiella sp.,Oulodus shiqianensis,Oulodus sp.,Pseudooneotodus beckmanni,Distomodus sp.。根据牙形刺的产出情况,可识别出两个牙形刺带。秀山组顶部灰岩可归入Pterospathodus eopennatus超带的P.eopennatus ssp.n.2带。需要指出的是,秀山组最顶部样品发现了P.a.cf.angulatus,因此,秀山组顶部的时代延伸至Pterospathodus celloni超带的P.a.angulatus带,时代为兰多维列世特列奇早-中期。     

湖北宜昌黄花场下奥陶统弗洛阶Oepikodus evae带精细地层划分对比

全球中奥陶统及奥陶系第三个阶、大坪阶的界线层型剖面和点位(GSSP)确定后,文中作者对宜昌黄花场剖面弗洛阶上部Oepikodus evae带牙形刺生物地层序列进行再研究。通过对红花园组上部-大湾组下段一些关键层位中牙形刺样品的补充采集和牙形刺标本的系统检查,研究牙形刺属种垂向分布特征,修订一些牙形刺属种的首现层位。讨论Stolodus stola,Lundodus gladiatus,Oepikodus intermedius,Oepikodus communis,Trape-zognathus diprion,Baltonioduscf.B.triangularis等重要的多分子器官种分类问题。识别出Baltoniodus trian-gularis(Lindstr m)的先驱:Baltonioduscf.B.triangularis(Lindstr m)与Baltoniodus triangularis(Lindstr m)的区别。描述新建化石带属种,包括Baltoniodus triangularis(Lindstr m)。认为Oepikodus evae带上部时限范围内并未见Oepikodus evae分子,所以与国际地层指南(2000)关于间隔带和延限带的含义相悖。据此,厘定了原Oepikodus evae带的含义。并以牙形刺带分子首现为标志,将大湾组下段自下而上划分为3个间隔带,分别为:O.evae带,Trapezognathus diprion带和Baltonioduscf.B.triangularis带。其中Trapezognathus diprion带和Baltonioduscf.B.triangularis带为新建化石带。     

蒙古南戈壁泥盆纪(埃姆斯期-艾菲尔期)牙形刺

在蒙古南戈壁地区丘隆组(Chuluun Formation)和查干卡勒噶组(Tsagaankhaalga Formation)发现的牙形刺证明丘隆组的时代为晚埃姆斯期,查干卡勒噶组的时代为早艾菲尔期,与以前大化石所确定的时代一致。但艾菲尔阶的底界暂时还无法准确确定,蒙古南戈壁地区艾菲尔阶牙形刺动物群与同期北美的牙形刺动物群相似．但也包含一些地方性分子,蒙古海相艾菲尔期的沉积可能不完整。本文描述了2个命名新种．Caudicriodus delicatus sp. nov. , C. paraculicellus sp. nov. ,一个未命名新种,Caudicriodus sp. nov. A和一个新亚种,Cuadicriodus angustus cauda subsp, nov.。     

豫淮盆地太原组顶部斯威特刺(Sweetognathus)种的分类修正及其地层意义

作者通过对豫淮盆地太原组顶部灰岩中牙形刺的研究,发现Sweetognathus动物群较为丰富,计4属10种,含一个新种:Sweetognathuspraeiranicussp.nov.。其中以Sweetognathus最为繁盛,并对其种的分类进行了修正,将原定的Sweetognathusinornatus的大部分标本归入Sweetognathusmerrilli。根据新的分类,建立了两个组合带,即Sweetognathusmerrilli带和Sweetognathuswhitei-Xuzhougnathusmonoridgosus带。通过与国内外相当牙形刺组合带(带)的对比,初步认为该动物群属中、晚萨克马尔到早亚丁斯克期。     

四川广元旺苍罐子坝剖面长兴组与大隆组牙形刺及其生物地层研究

系统研究了四川东北部广元市旺苍县罐子坝剖面长兴组和大隆组的牙形刺动物群,鉴定2属21个种(包括2个相似种,1个亲缘种和1个未定种)。划分出4个牙形刺带,从下至上依次为Clarkina orientalis带,C.wangi带,C.changxingensis-C.deflecta带和C.meishanensis带。以C.wangi的首现层面为标志比较精确地确定了该剖面长兴阶的底界,并将该剖面牙形刺序列与浙江长兴煤山经典剖面进行了较为精确的对比。     

Famennian conodonts from the Hongguleleng Formation at the Bulongguoer stratotype section,western Junggar,Northwest China

The Hongguleleng Formation, the highest and most important Devonian marine carbonate horizon in western Junggar, contains an endemic shallow-water Icriodus–Polygnathus conodont fauna with rare palmatolepids and other genera. The conodont faunas from the Bulongguoer and the Wulankeshun sections are similar, with high abundance (about 40%) of endemic taxa (13 taxa), indicating isolation of the Junggar Basin during early Famennian. Non-endemic species in the faunas suggest that the Lower Member of the Hongguleleng Formation is assignable to the Pa. rhomboidea Zone to the Pa. marginifera marginifera Zone of early Famennian, not including the Frasnian–Famennian boundary. The Upper Member may be of late Famennian–early Tournaisian in age on the basis of our preliminary faunal analysis.     

吉林李家窑范家屯组中的二叠纪北温带牙形刺动物群

在吉林省九台市三台乡李家窑附近的二叠系范家屯组中采到一特殊的牙形刺动物群。多数牙形刺都较小,锯齿发育,齿脊细齿不愈合。由一些未见报道的新分子组成。本文描述了４个新种（Ｍｅｓｏｇｏｎｄｏｌｅｌｌａ ｃｈａｎｇｃｕｎｅｎｓｉｓ ｓｐ．ｎｏｖ．,Ｍ．ｊｉｌｉｎｅｎｓｉｓ ｓｐ．ｎｏｖ．,Ｍ．ｍｕｌｔｉｓｅｒｒａｔａ ｓｐ．ｎｏｖ．,Ｍ．ｐｓｅｕｄｏａｌｔｕｄａｅｎｓｉｓ ｓｐ．ｎｏｖ．）和３个未命名新种     

Sedimentary and faunal changes across the frasnian/famennian boundary in the canning basin of Western Australia

The Canning Basin of northwestern Australia is a key area for understanding global changes at the “Kellwasser Events” and the Frasnian‐Famennian boundary. Frasnian stromatoporoid‐coral‐cyanobacterial reef platforms stretched out for enormous distances along the palaeoshelf but in the early Famennian they were completely replaced by cyanobacterial reef platforms. An iridium anomaly in the sequence was formerly believed to be at or close to the boundary and was interpreted as possible evidence for an asteroid impact. Recent field work and detailed biostratigraphy in the area east and southeast of Fitzroy Crossing has given dating relevant to the timing and extent of sea level changes, hypoxic incursions and reef backstepping. Goniatites and conodonts provide correlations with the international biostratigraphy.

In the Horse Spring area the stage boundary falls within the Virgin Hills Formation which normally has a rich pelagic goniatite, nautiloid and conodont fauna. In the latest Frasnian (Zone 13 of Klapper 1989; regional Sphaeromanticoceras lindneri Zone) large allochthonous reef blocks moved downslope into the open marine basin. A diverse gastropod fauna is associated with the last atrypid brachiopods. The faunal record at the immediate boundary is obscured by dolomitisation but manticoceratid goniatites range into this level. There is no evidence for the organic‐rich dark Kellwasser limestone facies.

In the McWhae Ridge area two Frasnian goniatite horizons with Beloceras trilobites and the giant Mamicoceras guppyi and Sphaeromanticoceras lindneri transgress over the reef slope. Stromatolitic cyanobacterial beds mark condensations. Again there is no trace of the oxygen‐depleted Kellwasser facies. However, as at Horse Spring, manticoceratids persisted into dolomitic marker beds that have no other preserved macrofauna. The iridium anomaly associated with Frutexites postdates the Frasnian‐Famennian boundary and was formed by cyanobacterial concentration.     

Frasnian conodonts from the eastern Russian Platform

The Frasnian conodonts of the eastern Russian Platform from the Timan in the north to the Orenburg Region in the south are characterized in detail both paleontologically and stratigraphically. This paper is a continuation of a previously published monograph (Ovnatanova and Kononova, 2001) on the Frasnian conodonts of the shallow-water strata of the central Russian Platform, where polygnathids prevail. The conodont assemblages from the shallow-water and basinal sections of the Volga-Ural province and Southern Timan are analyzed. Problems and difficulties associated with the correlation of the shallow-water and deep-water Frasnian sections of the eastern Russian Platform and their correlation with the existing zonal conodont scales are discussed. The correlation between the deep-water Mendym deposits and the shallow-water Rechitza and Voronezh strata of the Volga-Ural province with the Vetlasyan and Sirachoi deposits of the Southern Timan is substantiated. The stratotype of the Semiluki Regional Stage of the Central Devonian Field contains equivalents of the lowermost beds of the Domanik Formation (unit 1), based entirely on polygnathids. Phylogenetic reconstructions for Palmatolepis and Polygnathus are suggested based on the ontogenetic series for some species of these two genera and the presence of transitional forms between some of the species. Based on these phylogenetic reconstructions, conodont zonal scales for the shallow-water and deep-water sections of the Frasnian Stage of the Russian Platform are proposed and their correlation with the existing conodont zonal scales is also adduced. Some aspects of biofacies control are considered based on the distribution of conodonts in the sections studied. In Systematic Paleontology, 91 conodont species of the genera Ancyrodella, Mesotaxis, Palmatolepis, and Polygnathus are described, including the new species Palmatolepis menneri, P. kaledai, P. acutangularis, and Polygnathus reitlingerae. DEDICATION To the blessed memory of the outstanding geologist Vladimir Vladimirovich Menner     

MICROVERTEBRATE BIOSTRATIGRAPHY OF UPPER DEVONIAN (FRASNIAN) CARBONATE ROCKS IN THE CANNING AND CARNARVON BASINS OF WESTERN AUSTRALIA

Abstract:  Shark teeth and scales from the Gneudna Formation type section, Carnarvon Basin are rare, but they represent a diverse fauna as well as being the first Frasnian chondrichthyan remains found in Western Australia. In contrast numerous shark teeth and scales have been obtained from coeval sections in the Canning Basin. Teeth referred to as Phoebodus bifurcatus, Phoebodus fastigatus, Phoebodus cf. fastigatus, Phoebodus latus , Phoebodus sp. C and Protacrodus sp. 1 are described from two Canning Basin localities: Horse Spring Range and McIntyre Knolls. The phoebodont species described here have a global distribution and can, therefore, be placed within the standard Frasnian phoebodont zonation. The shark remains from the Gneudna Formation type section include one new genus Emerikodus ektrapelus gen. et sp. nov., described along with Helodontidae indet and Elasmobranchii gen. et sp. indet. It is proposed that the faunal differences observed between the localities are a result of environmental differences. In addition to shark remains, scales from the thelodont Australolepis seddoni were also recovered from each locality. The well-dated Zone 6–10 conodont faunas at Horse Spring constrain the range of A. seddoni with which they occur, and importantly indicate that the Gneudna Formation type section is wholly Frasnian.     

新疆准噶尔盆地西北缘洪古勒楞组时代的新认识

标准地点的洪古勒楞组长期以来一直被认为是准噶尔盆地西北缘,乃至整个新疆北部以海相为主的层位上最高的晚泥盆世晚期的沉积。然而牙形类和微体脊椎动物的研究证实,该组跨弗拉阶－法门阶（Ｆ－Ｆ）界线,它至少包括了一小部分弗拉晚期和主体部分为法门早期的沉积。两个牙形类带ＬａｔｅｒｈｅｎａｎａＺｏｎｅ和ＭｉｄｄｌｅｃｒｅｐｉｄａＺｏｎｅ被识别出,它们在Ｆ－Ｆ界线附近,并分别在大绝灭事件（在ｌｉｎｇｕｉｆｏｒｍｉｓＺｏｎｅ内）之前后。另一个层位更高的牙形类带ＬａｔｅｃｒｅｐｉｄａＺｏｎｅ也可能存在。同时还证实,长期以来一直被认为“正常层序”的标准地点的洪古勒楞组的层序是倒转的。由此,含植物大化石Ｌｅｐｔｏｐｈｌｏｅｕｍｒｈｏｂｉｃｕｍ等的陆相层在下（此层可能归于该组下伏的朱鲁木特组）,而产牙形类和其它丰富的无脊椎动物的海相层在上。按牙形类年代带的年代地层时间表（Ｆｏｒｄｈａｍ,１９９２）,该组距今约３６８－３６４百万年。至于它和邻近地区和布克河组的关系,由于后者发现了更高层位的牙形类,两者还不能完全对比。     

滇东北昭通箐门的泥盆纪珊瑚(三)———晚泥盆世早期的四射珊瑚*

云南昭通市东北郊箐门出露了一条比较完整的泥盆纪浅海相地层剖面,下、中、上三个统均有出露,浅海底栖生物化石丰富,研究基础也相对较好,是研究滇东北地区泥盆纪地层的一条理想剖面。由于地层剖面比较长,沉积厚度比较大,此前笔者已经描述了早泥盆世和中泥盆世的四射珊瑚,本文继续描述该剖面的晚泥盆世的地层及其四射珊瑚,讨论它们的地层划分、珊瑚群对比与古生物地理。昭通箐门剖面的上泥盆统主要由白云质灰岩组成,与滇东其它地方的宰格组的岩性基本一致。过去由于宰格组的化石稀少,其地层时代难于确定,因此,昭通箐门宰格组底部珊瑚化石的发现具有重要意义。文中首次描述了箐门放羊冲剖面宰格组底部的四射珊瑚5属7种,包括Disphyllum catenatum,Hexagonaria davidsoni,H.magna,Sinodisphyllum simplex,S.variabile,Temnophyllum poshiense和Truncicarinulum temeniophylloides等,总体显示了晚泥盆世弗拉早期(early Frasnian)的特点。它除了与华南各省的珊瑚群有密切关系外,也与加拿大西部的分子有一定的联系。     

Conodonts from the Permian Jisu Honguer (Zhesi) Formation of Inner Mongolia, China

The Jisu Honguer Formation (“Zhesi Formation”) is a North China marine carbonate unit of Permian age containing a mixed fauna of Tethyan, Boreal and endemic elements. The age of the Jisu Honguer Formation has been thought to range from Artinskian to Kazanian based on previous studies using mostly benthic macrofossils. A typical Mesogondolella aserrata conodont fauna is reported from the lower part of the upper member of the Jisu Honguer Formation in the Ulanqub District, Zhesi area, of northern China. The fauna indicates a Middle Permian (Guadalupian) age, most likely late Wordian to early Capitanian, for the strata yielding conodonts. The whole Jisu Honguer Formation could be assigned to the Wordian to early Capitanian age. The overlying Yihewusu Formation is probably of Capitanian age. There are no Wuchiapingian marine deposits in the Zhesi area. As characteristic of the open sea Guadalupian conodont faunas of the Tethys, smooth Mesogondolella dominate the fauna. The view that all Tethyan Guadalupian conodont faunas consist exclusively of serrated Mesogondolella cannot be confirmed. Three new species are described: Mesogondolella neoprolongata C. - y. Wang, Mesogondolella mandulaensis C. - y. Wang and Wardlawella jisuensis C. - y. Wang.     

Environmentally controlled phyletic evolution, blindness and extinction in Late Devonian tropidocoryphine trilobites

The Middle and Upper Devonian carbonate succession of the Montagne Noire, Southern France has been precisely zoned by an unbroken sequence of conodont zones. Stratigraphic control is excellent, and has allowed evolutionary changes in tropidocoryphine trilobites, which occur throughout the succession, to be directly established. The tropidocoryphine had been a stable group for some 40 million years prior to the Middle Givetian. In their last few million years, however, they underwent rapid evolution and exhibit some striking transformations of the cephalon and the regression and virtual disappearance of the eye within a relatively short space of time. They also show a marked diminution in size, and lose their original relief so that the glabella becomes virtually flush with the surface. There are two separate lineages, both of which show eye-reduction and subsequent blindness. In the earlier lineage Tropidocoryphe (Longicoryphe)-Erbenicoryphe, the main features of the ancestral rootstock are conserved and the stable, strongly divergent anterior sutural pattern of the cephalon remains the same. The eye, however, became reduced to a slightly convex surface lacking lenses, only indistinctly defined. Erbenicorphe is confined to well-oxygenated facies, and probably lived as a shallow burrower within the sediment; it became extinct in the early Frasnian. The second lineage T. (Longicoryphe)-Pterocoryphe- Pteroparia shows a remarkable backward migration of the suture, which progressively swings posteriorly in successive species spanning four Frasnian conodont zones (about three million years duration). At the same time the eye progressively degenerates so that the last forms are blind. Sutural migration and eye reduction are not genetically linked, however; the unusual form of the cephalon and suture probably resulted from an adaptation to the euxinic environment in which Pterocoryphe originated. The loss of the eye resulted from the adoption of an endobenthic habit in Pteroparia which descended from the ancestral Pterocoryphe but which had migrated to an oxygenated facies. Eye-reduction is therefore parallel in the two lineages, but superimposed upon a different original cephalic configuration. The last Pteroparia became extinct when the late Frasnian fauna became overwhelmed by the first pulse of the ‘Kellwasser Event’ (probably an anoxic overturn). The evolving characters, through virtually all observed steps, show progressive unidirectional change without sudden breaks or saltations. Such unidirectional evolution is an adaptive response to constant long-lasting environmental influences. □Evolution, Devonian trilobites, France, gradualism, eye reduction.