MORPHOLOGY AND ANATOMY OF FLORAL AND EXTRAFLORAL NECTARIES IN CAMPSIS (BIGNONIACEAE)

The genus Campsis (Bignoniaceae), with one New World and one Old World species, is unusual among temperate plants in having five distinct nectary sites. Multiple nectaries occur at all four of the extrafloral sites (petiole, calyx, corolla, fruit), representing an advanced strategy for ant attraction. The morphology and anatomy of the extrafloral nectaries in both species are uniform for the petioles, calyces, and young fruits; those on the outer corolla lobes are of slightly different forms. The generalized structure consists of one layer of basal cells, and a one- to two-layered secretory cup. Because of their small size, there is no vascular tissue in them. The large, vascularized (phloem only) floral nectary is an annular structure subtending the ovary.