Pleiotropic effect of a pelargonidin-hydroxylation gene in Silene dioica?

In petals of Silene dioica a gene P has been identified, which controls the 3′-hydroxylation of the B-ring of pelargonidin to cyanidin. Another gene Ac controls the acylation of the terminal sugar at the 3-position of anthocyanin 3-rhamnosylglucoside-5-glucosides. In p/p plants the bound acyl group is p-coumaric acid; in P/P plants, however, it is caffeic acid. Gene P seems to exert a pleiotropic effect: it not only controls the hydroxylation of the B-ring of pelargonidin but also that of the acyl group.