1. Department of Biological Sciences, Vanderbilt University, Nashville, TN 37235, USA;2. Department of Synthetic Biology and Bioenergy, The J. Craig Venter Institute, Rockville, MD 20850, USA;3. Department of Electrical Engineering and Biological Science, Waseda University, TWIns, 2-2 Wakamatsu-cho, Shinjuku, Tokyo 162-8480, Japan