1. Department of Cognitive Neurology, Hertie Institute for Clinical Brain Research, University of Tübingen, Hoppe-Seyler-Strasse 3, 72076 Tübingen, Germany;2. Department of Neurodegeneration, Hertie Institute for Clinical Brain Research, University of Tübingen, Hoppe-Seyler-Strasse 3, 72076 Tübingen, Germany;3. German Centre for Neurodegenerative Diseases (DZNE), University of Tübingen, Otfried-Müller-Strasse 27, 72076 Tübingen, Germany