1. Department of Neuroscience, Physiology and Pharmacology, University College London, London WC1E 6BT, UK;2. Institute of Cognitive Neuroscience, University College London, London WC1N 3AR, UK;3. Wolfson CARD, King’s College London, London SE1 1UL, UK;4. Nuffield Department of Clinical Neurosciences, University of Oxford, Oxford OX3 9DU, UK