Clustered trilobite assemblages in the Middle Devonian Hamilton Group

Clusters of well-preserved trilobites occur throughout lower and middle Paleozoic strata. This phenomenon, however, remains poorly understood. Several thin, regionally persistent beds in the Middle Devonian (Givetian) Hamilton Group of New York State display taphonomic and sedimentary features which indicate rapid burial by storm-generated clouds of fine-grained sediment. Furthermore, components in these units display little or no post-mortem disturbance and, therefore, reflect in faunal associations. These beds contain two types of clustered assemblages of the trilobites Phacops (Green), Greenops boothi (Green) and Dechenella rowi (Green). Body clusters comprise dense assemblages of three or more complete trilobite corpses. Moult clusters , in contrast, are composed of three more moult ensembles; i.e., articulated thoracopygidia in close proximity to corresponding cephala. Clusters are species-segregated and contain only a small portion of the total size range exhibited by trilobite species. Moreover, body clusters and moult clusters are morphometrically undifferentiated indicating that these two cluster modes represent related phenomena. Analogy with modern marine arthropods suggests that trilobites assembled into monospecific, age-segregated clusters and moulted prior to en masse copulation.