The Papuan tortricine genus Heterochorista and the systematic position of the Sparganothini (Lepidoptera, Tortricidae)

ABSTRACT .
The previously monotypic Papuan genus Heterochorista Diakonoff, 1952 is revised and twenty-one species are recognized. Nikolaia Diakonoff, 1953 is synonymized, syn.n. with Heterochorista. The following six species are placed in new combination in Heterochorista: Nikolaia melanopsygma Diakonoff, N.inumbrata Diakonoff, N.aperta Diakonoff, Isochorista papuana Diakonoff, I.polysperma Diakonoff and Dicellitis chrysonetha Diakonoff. Thirteen new species are added to Heterochorista: acomata sp.n., rostrata sp.n., trivialis sp.n., ornata sp.n., prisca sp.n., punctulata sp.n., rufulimaculata sp.n., classeyiana sp.n., spinosa sp.n., signata sp.n., fulgens sp.n., nitida sp.n. and aura sp.n. A checklist and a key to species are provided, and adults and male and female genitalia are described and illustrated. The phylogeny of Heterochorista is outlined on the basis of a cladistic character analysis and the genus is referred to the base of the Sparganothini. The apomorphies characterizing this tribe are reviewed with regard to their phylogenetic significance and the relationship between Sparganothini, Archipini and Atterini is briefly discussed.