Two new genera of Laboulbeniales allied to Zodiomyces

Two new genera of Laboulbeniales (Ascomycota) allied to Zodiomyces (Zodiomycetoideae) are described. These are Capillistichus with the single species C. tenellus, parasitic on Laccobius spp. (Coleoptera, Hydrophilidae), and Scepastocarpus with the single species S. peritheciiformis, parasitic on Hydrochus spp. (Coleoptera, Hydrophilidae). Both genera and species are from Spanish collections. Perithecial characters appear to be similar in the two new genera and in the related Zodiomyces, becoming a diagnostic character of the group (subfamily Zodiomycetoideae). Receptacular characters form the basis of separation into genera. To complete the study of all the taxa included in the subfamily, specimens and type slides of all three species of Zodiomyces have been reviewed.