The Enidae of Timor (Stylommatophora: Orthurethra)

The systematic classification of the Enidae of Timor is reviewed by means of comparative morphology and molecular phylogenetics using recently collected land snail samples from Timor-Leste (‘East Timor’). Features of the renal and genital anatomy as well as analyses of nuclear 28S rRNA sequences confirm the placement of the species in the Enidae. Generic combinations employed by earlier authors are discussed. The name Coccoderma Möllendorff, 1901 used for enids from the Indo-Australian Archipelago is pre-occupied by Coccoderma Zittel, 1887 and therefore not available. The assignment of species from Timor to Pseudonapaeus Westerlund, 1887, which is otherwise known from central Asia, is also rejected. Instead, the Enidae from Timor are here affiliated with Apoecus Kobelt, 1902, a name originally introduced for the Papuan species Buliminus colonus Möllendorff, 1895. While Apoecus samples from Timor-Leste form three distinct mitochondrial clades, only two groups can be reliably distinguished by their morphology. One of these groups represents the previously named species Apoecus apertus (Martens, 1863). This species is found throughout lowland Timor occurring at altitudes of up to 600?m. The second species has been found in the Ramelau Mountains at altitudes above 1300?m, and is described as Apoecus ramelauensis n. sp.

http://zoobank.org/urn:lsid:zoobank.org:pub:3A8781B6-6EAC-4C58-8DB3-63741B03B43C