The first representative of the suborder Mesotitanina from the Paleozoic and notes on the system and evolution of the order Titanoptera (Insecta: Polyneoptera)

A new subfamily, genus, and species (Deinotitaninae subfam. nov., Deinotitan orenburgensis gen. et sp. nov.) of the family Mesotitanidae (Titanoptera) are described from the Severodvinian (Upper Permian) of the Orenburg Region. This family belongs to the suborder Mesotitanina; both are recorded in the Paleozoic for the first time. The present finding and unpublished data suggest that the Mesozoic stage of the evolution of the superorder Orthopteroidea might start in the Late Permian. Taxonomy and early evolution of the order Titanoptera are discussed. Minititan nom. nov. is proposed for Microtitan Gorochov, 2003 (nec Microtitan Granger et Gregory, 1943).