Carminator, a new genus of Megalyridae (Hymenoptera) from the Oriental and Australian regions, with a commentary on the definition of the family

The taxonomic history of the family Megalyridae is reviewed. Carminator gen.n. from the Oriental and Australian regions is described and illustrated. Four new species are included in the genus: C.affinis, Cater, C.cavus and C.nooni. A key to the species is provided.
Twelve characters are evaluated for their phylogenetic significance, including those which historically have been regarded as important for the definition of the family. Carminator is provisionally assigned to the subfamily Dinapsinae, based on the reduction of Rs in the hindwing. The family Megalyridae is regarded as monophyletic based on their unique pattern of reduced hindwing venation and position of the 'pronotalrsquo; spiracle. The mesozoic Cleistogastrinae are excluded from Megalyridae. The cretaceous genus Maimetsha is reclassified as a dinapsine megalyrid.