Spores and the middle-upper Devonian boundary

The stratigraphic levels most favoured for the Middle-Upper Devonian boundary fall approximately within the range of the ammonoid Pharciceras lunulicosta Zone, i.e. from the Middle varcus Subzone to the base of the Lower asymmetricus Zone of the conodont scale. Spore data that are potentially useful for recognition of the boundary within this range have been correlated with conodont zones in marine facies in the Boulonnais region of France. A vast amount of information on spores from Middle-Upper Devonian boundary strata has accumulated in the European U.S.S.R., where the boundary is taken at a somewhat lower level. Late Givetian and early Frasnian continental strata of Melville Island in the Canadian Arctic contain species present in the Boulonnais or the European U.S.S.R., as well as species common to both regions. Diatomozonotriletes spp., Rhabdosporites langii, Samarisporites triangulatus, Contagisporites optivus, Archaeoperisaccus timanicus, Chelinospora concinna and Ancyrospora langii, among others, may be useful for correlating the boundary as eventually defined. The stratigraphic ranges of most of these taxa show only limited agreement interregionally at present, probably owing at least in part to problems of spore nomenclature and taxonomy, and an insufficiency of spore reference sequences keyed to faunal zones. Nevertheless, individual species of spores, and especially assemblages of species, have much potential for delimiting and correlating the Middle-Upper Devonian boundary in both marine and continental facies.