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Cenomanian cheilostome bryozoans from Devon,England
Institution:1. Department of Earth Sciences, Natural History Museum, Cromwell Road, SW7 5BD London, United Kingdom;2. Senckenberg Forschungsinstitute und Naturmuseen, Sektion Marine Evertebraten III (Bryozoologie), 25, Senckenberganlage, 60325 Frankfurt am Main, Germany;1. Senckenberg German Entomological Institute, Müncheberg D-15374, Germany;2. Zoology, Institute for Biology, Faculty of Natural Sciences I, Martin-Luther-University Halle-Wittenberg, Halle D-06099, Germany;3. Department of Biogeography, Trier University, Trier D-54286, Germany;1. Department of Earth and Environmental Sciences, Sec. Geophysics, Ludwig-Maximilians Munich University, Theresienstrasse 41, 80333 Munich, Germany;2. ROCEEH Research Centre “The Role of Culture in Early Expansions of Humans”, Heidelberg Academy of Sciences, Senckenberg Research Institute, Senckenberganlage 25, 60325 Frankfurt am Main, Germany;3. Armenian National Academy of Sciences, Institut of Botany, Avan-63, Yerevan 375063, Armenia;4. Department of Earth Sciences, Utrecht University, Budapestlaan 17, 3584 CD Utrecht, The Netherlands;1. Institut für Geologie, Universität Hamburg, Bundesstr. 55, 20146 Hamburg, Germany;2. Kuban State University, Kazan Federal University, Aphipskij, Post box demand, 353235 Krasnodar, Russia;3. Département de géologie, université de Liège, bâtiment B18, Sart-Tilman, 4000 Liège, Belgium;4. Institut royal des sciences naturelles de Belgique, DO terre et histoire de la vie, 29, rue Vautier, 1000 Bruxelles, Belgium;1. Instituto de Geología, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, México, D.F., Mexico;2. Departamento de Ingeniería Geológica, Facultad de Ingeniería, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, México, D.F., Mexico;3. Sovereign Pharmaceuticals, 7590 Sand St., Fort Worth, 76118, TX, United States;4. Département des Sciences de la Terre, Université de Nice-Sophia Antipolis, 28 Avenue Valrose F-06100 Nice, France;5. Facultad de Estudios Superiores Iztacala, UNAM, Tlalnepantla, Estado de México, 54070, Mexico
Abstract:The Cenomanian witnessed a spectacular evolutionary radiation of cheilostome bryozoans, both in terms of species diversity and morphological disparity. However, Cenomanian cheilostome faunas are inadequately known. Twelve species of cheilostome bryozoans are here described from the Cenomanian Beer Head Limestone Formation of SE Devon, England, a nearshore facies of limestones and sands. Two of the cheilostome species are new – Wilbertopora manubriformis sp. nov. and Foratella cervisia sp. nov. – and five cannot be identified beyond genus level. Syntypes of Foratella forata (d’Orbigny, 1853), the Senonian type species of Foratella Canu, 1900, are illustrated using SEM and a lectotype is chosen. All of the species present are neocheilostomes, which were larval brooders. Compared with the non-brooding malacostegans that dominate pre-Cenomanian faunas, most species have avicularia and extensive frontal walls, features probably adaptive against small predators.
Keywords:Bryozoa  Cheilostomata  Taxonomy  New taxa  Cretaceous  Cenomanian  England  Bryozoa  Cheilostomata  Taxinomie  Nouveaux taxons  Crétacé  Cénomanien  Angleterre
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