Affiliation: | 1. Department of Biological Sciences, University of Idaho, Moscow, ID, USA;2. Parc National des écrins, F‐05000, France;3. Laboratoire d'Ecologie Alpine, CNRS ‐ Université Grenoble Alpes, UMR 5553, Grenoble, France;4. Institute of Geological Sciences, University of Bern, Bern, Switzerland |