Affiliation: | 1. Department of Wildlife Biology, Forestry and Forest Products Research Institute, 1 Matsunosato, Tsukuba, Ibaraki, 305-8687, Japan 2. Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi, Yaeyama, Okinawa, 907-1541, Japan 3. Graduate School of Agricultural Science, Tohoku University, 232-3 Yomogida, Naruko-onsen, Osaki, Miyagi, 989-6711, Japan 4. Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa, 903-0213, Japan 5. Iriomote Forest Ecosystem Conservation Center, Kyushu Regional Forest Office, 55-4 Tonoshiro, Ishigaki, Okinawa, 907-0004, Japan
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