山茱萸科花粉形态研究

本文应用光学显微镜与扫描电子显微镜观察了山茱萸科Cornaceae4属(按照Wangerin1910年的概念)27种花粉形态,结果如下: 1.根据花粉特征,支持把青荚叶属Helwingia,桃叶珊瑚属Aucuba,鞘柄木属Torricellia三属分别独立成科的观点。 2.梾木属Cornus可明显区分为两种类型,即梾木型和四照花型;梾木型还可再分为三个亚类型.即灯台树亚型,长圆叶梾木亚型和梾木亚型。 5.支持四照花属应从梾木属中分出而独立成属;而在四照花属Dendronbenhamia下设立北美四照花亚属Subg. Apocarpea和四照花亚属Subg. Syncarpea;在梾木属下设立梾木亚属Subg. kraniopsis, 灯台树亚属Subg.Masomora等亚属的观点。 4.建立了—新亚属——长圆叶梾木亚属。

STUDIES ON POLLEN MORPHOLOGY OF CORNACEAE

The present paper describes the pollen morphology of 27 species representing 4 genera of Cornaceae (Wangerin's 1910 system), the pollen grains were all examined under light, scanning electron microscope. The results are as follows:Helwingia, Aucuba, Torricellia have respectively their pollen characteristics, no similarities with the other genera of Cornaceae. Systems of J. Decaisne(1836), A. de Condolle(1868) and Takhtajan(1969) treated Helwingia, Aucuba and Torricellia as three separate families: i. e. Helwingiaceae,Aucubaceae and Torricelliaceae.The results of the study support their systems.Based on the characters of the structure of tectum, pollen grains of the genus Cornus L. may be divided into two types : Cornus-type and Dendronbenthamia-type. The former includes the examined species of Cornus (in T. Nakai's 1909,J. Hutchinson's 1942 systems) which have pollen with mark perforate tectum; the latter includes the examined species of Dendronbenthamia, Macrocarpium (in T. Nakai's 1909,J. Hutchins-on's 1942 systems) which have pollen with no perforate tectum.According to the feature of muri, the pollen grains of Cornus-type can be divided into three subtypes: i. e. Alternifolia-subtype,Oblongasubtype and Cornus-subtype.Through analysis of pollen characteristics and plant morphorlogy of Cornus and Dendronbenthamia, the paper agrees with that Dendrobenthamia and Cornus should be different genera, and with that subg. Kraniopsis Raf., subg. Mesomora Raf., subg Oblongania (Wanger.)K. Y. Wang, subg. nov. should be established in Cornus and subg. Apocarpea Nakai, subg. Syncarpea Nakai in Dendronbenthamia.