Genetic identity in subdivided populations. I. Two equal-sized subpopulations

Moments of the steady state frequency spectrum (probabilities of identity of samples of genes) are obtained for a subdivided population by using standard recursive identity by state calculations. These moments are used to obtain variances for some measures of genetic identity, including Nei's normalized genetic identity (I) and genetic distance (?log_eI). The results are compared with those obtained from the corresponding undivided population theory, including adjustments to the effective number to try to account for subdivision. Undivided population approximations based on effective number are surprisingly accurate, regardless of the migration rate, when sampling exclusively from one subpopulation.