Institution: | 1 Research Department of Genetics Evolution and Environment, University College London, Darwin Building, London WC1E 6BT, UK;2 Department of Biology, University of Addis Ababa, PO Box 32597, Ethiopia;3 Department of Science, Systems and Models, Roskilde University, 4000 Roskilde, Denmark;4 Henry Stewart Group, 28/30 Little Russell Street, London WC1A 2HN, UK |