Archaeological evidence of former occurrence and changes in fishes, amphibians, birds, mammals and molluscs in the Wadden Sea area

Animal remains are well preserved in archaeological sites, especially the terp sites, of the Wadden Sea area of Denmark, Germany and The Netherlands. Here, we provide an overview on the wild mammals, birds, fishes, amphibians and molluscs found in coastal sites dating from 2700 to 2600 B.C. and 700 B.C. to A.D. 1600. Coastal people used a variety of animal species for food and other purposes. Hunting, fowling, fishing and agriculture did not have much influence on wild stocks in the period from the late Bronze Age/early Iron Age until the late Middle Ages. However, large changes to the landscape were made in the late Middle Ages by diking and damming. As a result, some species such as the northern vole (Microtus oeconomus) and the natterjack toad (Bufo calamita) disappeared from the area except for some dune districts on the islands, and others became rare, such as the grey seal (Halichoerus grypus) and the lagoon cockle (Cerastoderma lamarcki). New habitats arose for birds of dry meadows and fields, like lapwing (Vanellus vanellus) and black-tailed godwit (Limosa limosa). Sturgeon (Acipenser sturio) disappeared from the Wadden Sea within a few decades since A.D. 1890 due to the destruction of spawning grounds by damming and high exploitation pressure. Our findings are important for the ecological history of the region.