Department of Population Health and Reproduction, University of California, Davis, CA, 95616 USA;2. Genetics Research Laboratory, California Department of Fish and Wildlife, Rancho Cordova, CA, 95670 USA;3. Arizona Game and Fish Department, Phoenix, AZ, 85086 USA;4. Nevada Department of Wildlife, Reno, NV, 89511 USA;5. Department of Fisheries and Wildlife, Oregon State University, Corvallis, OR, 97331 USA;6. Center for Large Landscape Conservation, Bozeman, MT, 95771 USA;7. National Park Service, Grand Canyon National Park, Grand Canyon, AZ, 86023 USA;8. National Park Service, Biological Resources Division, Fort Collins, CO, 80525 USA;9. Wildlife Health Center, University of California, Davis, CA, 95616 USA;10. Veterinary Genetics Laboratory, University of California, Davis, CA, 95616 USA;11. Wildlife Genomics and Disease Ecology Laboratory, Department of Veterinary Sciences, University of Wyoming, Laramie, WY, 82070 USA