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Long‐proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae)
Authors:ANTONIO ARILLO  ENRIQUE PEÑALVER  RICARDO PÉREZ‐DE LA FUENTE  XAVIER DELCLÒS  JULIA CRISCIONE  PHILLIP M BARDEN  MARK L RICCIO  DAVID A GRIMALDI
Institution:1. Departamento de Zoología y Antropología Física, Facultad de Biología, Universidad Complutense, Madrid, Spain;2. Museo Geominero, Instituto Geológico y Minero de Espa?a, Madrid, Spain;3. Museum of Comparative Zoology, Harvard University, Cambridge, MA, U.S.A.;4. Departament d'Estratigrafia, Paleontologia i Geociències Marines, Facultat de Geologia, Universitat de Barcelona, Barcelona, Spain;5. Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, U.S.A.;6. Institute of Biotechnology, Cornell University, Ithaca, NY, U.S.A.
Abstract:The monophyletic family Zhangsolvidae comprises stout‐bodied brachyceran flies with a long proboscis and occurring only in the Cretaceous, originally known in shale from the Early Cretaceous Laiyang Formation (Fm.) in China (Zhangsolva Nagatomi & Yang), subsequently from limestones of the Early Cretaceous Crato Fm. of Brazil. Cratomyoides Wilkommen is synonymized with Cratomyia Mazzarolo & Amorim, both from the Crato Fm.; Cratomyiidae is synonymized with Zhangsolvidae. Two genera and three species of Zhangsolvidae are described: Buccinatormyia magnifica Arillo, Peñalver & Pérez‐de la Fuente, gen. et sp.n. and B. soplaensis Arillo, Peñalver & Pérez‐de la Fuente, sp.n. , in Albian amber from Las Peñosas Fm. in Spain; and Linguatormyia teletacta Grimaldi, gen. et sp.n. , in Upper Albian–Lower Cenomanian amber from Hukawng Valley in Myanmar. Buccinatormyia soplaensis and Linguatormyia teletacta are unique among all Brachycera, extant or extinct, by their remarkably long, flagellate antennae, about 1.6× the body length in the latter species. A phylogenetic analysis of 52 morphological characters for 35 taxa is presented, 11 taxa being Cretaceous species, which supports placement of the family within Stratiomyomorpha, although not to any particular family within the infraorder. This published work has been registered in Zoobank, http://zoobank.org/urn:lsid:zoobank.org:pub:F32CF887‐7C37‐45D5‐BD6B‐135FE9B729A7 .
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