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First giraffid skulls (Bohlinia attica) from the late Miocene Maragheh fauna,Northwest Iran
Affiliation:1. Department of Vertebrate Paleontology, Natural History Museum of Los Angeles County, 900 Exposition Blvd., Los Angeles, CA 90007, USA;2. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China;3. State Key Laboratory of Lithospheric Evolution, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing 100029, China;4. Institutions of Earth Science, Chinese Academy of Sciences, Beijing 100029, China;5. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing 100049, China;6. Chinese Academy of Sciences Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China;7. Museum of Sonit Left Banner, Xilin Gol League, Inner Mongolia 011300, China
Abstract:An almost complete skull and a second partial skull of Bohlinia attica (Artiodactyla: Giraffidae) from the late Miocene of Maragheh in northwestern Iran is described along with a complete upper dentition from Samos in Greece. These specimens enrich what is known of this species. The skull has massive bent ossicones. The braincase is horizontal to the face in lateral view. The upper premolars have strongly curved styles. The metapodials of this species are elongate with a deep plantar trough. The braincase is short and the occipital does not extend caudally. The type material of Bohlinia is from Pikermi but this taxon is also known from localities in F.Y.R.O. Macedonia, Greece, Turkey, Bulgaria, Iraq, and Iran. The new specimens are similar to others except that the ossicone terminates in a knob and the palatine choanae are positioned caudally. This taxon along with Honanotherium, which is most similar but with shorter metapodials and simpler premolars, can be placed in the subfamily Bohlininae.
Keywords:Giraffidae  Turolian  Maragheh  Samos  Iran
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