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New material and revision of hipparions from the upper Miocene locality Ravin des Zouaves-5, RZO of Axios Valley (Macedonia,Greece)
Institution:1. Museum of Osaka University, 1-20 Machikaneyama-cho, Toyonaka, 560-0043 Osaka, Japan;2. Laboratory of Physical Anthropology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, 606-8502 Sakyo-ku, Kyoto, Japan;3. Faculty of Business Administration, Ryukoku University, 67 Tsukamoto-cho, Fukakusa, 612-8577 Fushimi-ku, Kyoto, Japan;4. Graduate School of Science and Engineering, Kagoshima University, 1-21-35 Korimoto, 890-0065 Kagoshima, Japan;1. Musée de la Géologie, Service Patrimoine Géologique, Office National des Mines, 24 Rue de l’Energie, 2035 La Charguia, Tunisia;2. School of GeoSciences, University of Edinburgh, James Hutton Road, Edinburgh, United Kingdom;1. Dipartimento di Scienze Chimiche e Geologiche, Università di Cagliari, Cittadella Universitaria (Blocco A), SS.554 bivio per Sestu 09042 Monserrato (CA), Italy;2. Dipartimento di Matematica e Geoscienze, Università di Trieste, via Weiss 2 34128 Trieste, Italy;1. Museum of Osaka University, 1-20 Machikaneyama-cho, Toyonaka, Osaka 560-0043, Japan;2. Laboratory of Physical Anthropology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto 606-8502, Japan;3. Faculty of Business Administration, Ryukoku University, 67 Tsukamoto-cho, Fukakusa, Fushimi-ku, Kyoto 612-8577, Japan;4. Graduate School of Science and Engineering, Kagoshima University, 1-21-35 Korimoto, Kagoshima 890-0065, Japan;1. University of Roma Tre, Department of Sciences, section of Geology, Largo S.L. Murialdo 1, 00186 Rome, Italy;2. Hungarian Natural History Museum, Department of Palaeontology and Geology, 1431 Budapest, Pf. 137, Hungary
Abstract:The reconsideration of the hipparions from the locality Ravin des Zouaves-5 (RZO) of the lower Axios Valley provides a better taxonomy, based on the revision of the old known collection (up to 1987), the inclusion of new discoveries and the bibliographic data of the last 30 years related to the study of the hipparions in Greece and the neighboring countries. Based on the cranial material and metapodials, five different taxa have been distinguished. New cranial material confirms the presence of the large-sized Hipparion proboscideum and the small-sized Hipparion macedonicum in the RZO fauna. In the new middle-sized hipparion sample from RZO two species have been identified, Hipparion philippus, and another form similar to Hipparion mediterraneum, which is attributed to Hipparion cf. mediterraneum. Cranial remains of a second small-sized hipparion are also recognized in the RZO fauna, which is attributed to Hipparion sithonis. The relationships of H. mediterraneum and H. proboscideum are discussed; a cladistic analysis suggests that they are sister taxa, sharing a common ancestor with the Hipparion moldavicum group, and a form similar to H. mediterraneum is a possible ancestor of H. proboscideum.
Keywords:Equidae  Turolian  Greece  Systematics  Biochronology  Evolution
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