REVISION OF THE GENUS NAPAEUS ALBERS, 1850 (GASTROPODA PULMONATA: ENIDAE). THE PROBLEM OF NAPAEUS (NAPAEINUS) NANODES (SHUTTLEWORTH, 1852) AND DESCRIPTION OF FIVE NEW SPECIES FROM ITS CONCHOLOGICAL GROUP

With this paper we initiate a taxonomic review of the genusNapaeus from the Canary Islands, indicating the statisticalmethodology used therein. We discuss the taxonomic problem ofNapaeus (Napaeinus) nanodes (Shuttleworth, 1852), designateits lectotype and describe five new species: Napaeus (Napaeinus)doliolum Henríquez, Napaeus (Napaeinus) bechi Alonso& Ibáñez, Napaeus (Napaeus) taguluchensisHenríquez, Napaeus (Napaeus) pygmaeus Ibáñez& Alonso and Napaeus (Napaeus) tagamichensis Henríquez,the first two from Tenerife; and the other three from La Gomera.Furthermore, we discuss the general relationships of the genusand its two sub-genera, whose original diagnoses are deficient;because of this, we think that in the future a broad reviewof the subfamily Eninae will be necessary in order to clarifythe actual position of this subfamily in the Canary Islands. (Received 11 March 1991; accepted 22 September 1992)