The European and eastern Nearctic fungus-gnats in the genus Ectrepesthoneura (Mycetophilidae)

Abstract. The validity and composition of Ectrepesthoneura Enderlein are discussed and it is contrasted with the related genus Tetragoneura Winnertz. The occurrence of mid-tibial sense organs in these genera and elsewhere in the subfamily Sciophilinae is considered. The European Ectrepesthoneura are revised and the secondary sexual characters of the male legs newly described. Two sgecies, E.pubescens (Zetterstedt ) = messaurensis Plassmann, syn.n. and E.colyeri sp.n., are added to the British list. The Nearctic species are discussed and the two eastern species dealt with fully: the male of E.bicolor (Coquillett) is newly recognized and Elaffooni sp.n. described. Some western Nearctic Tetragoneura probably belong correctly to Ectrepesthoneura but examination of types showed that T.quintana Cole, T.longicauda van Duzee and the eastern Nearctic T.pimpla Coquillett belong to Dziedzickia Johannsen in its present broad usage. The Palaearctic Ectrepesthoneura gracilis Edwards also probably has closer affinities with Dziedzickia than with its present assignment. The Japanese Ectrepesthoneura described by Sasakawa (1961) are excluded as they are probably Sciaridae.