Ontogeny of pistillate flowers and inflorescences in Nothofagus subgenus Lophozonia (Nothofagaceae)

  Cupule and pistillate inflorescence morphology is described in detail for the first time for a number of species in Nothofagus subgenus Lophozonia. The architecture and temporal appearance of prophylls, flowers, and cupule valves is dichasial; the cupule valves are third and higher order shoot systems. Comparative studies demonstrate that the dorsal rows of appendages on the cupule valves have a similar arrangement, and each row exhibits the same temporal pattern of development. This suggests that the cupule valves in all species have a modular construction and consist of rows of homologous appendages. This study clearly demonstrates the uniform groundplan of the pistillate inflorescence in this subgenus. Received March 14, 2002; accepted May 8, 2002 Published online: September 13, 2002