Redefinition of Ptycta Enderlein (Psocodea: "Psocoptera": Psocidae) and taxonomic revision of the Japanese species

The genera Ptycta Enderlein, 1925, and Copostigma Enderlein, 1903, are defined as a monophyletic complex based on the morphology of the male terminalia. Ptycta is redefined as those species of the Copostigma–Ptycta complex with forewing veins Rs+M fused for a short distance. Two new species of Ptycta from Japan are described, P. recava sp. nov. and P. johnsoni sp. nov., increasing the number of Japanese species to four, along with P. parvidentata Tsutsumi, 1964, and P. micromaculata Thornton, Lee, and Chui, 1972. Distributional information and illustrations of each species, and a key to Japanese species of Ptycta are included.