短柄五加大,小孢子发生和雌,雄配子体发育的研究

短柄五加花药５枚，每个花药四个花粉囊。小孢子母细胞减数分裂时，胞质分裂为同时型，产生正四面体形的四分体。花药壁由表皮、药室内壁、中层和绒毡层四层细胞组成，其发育类型为双子叶型。腺质绒毡层，其细胞为二核。三细胞型花粉。子房５室，每室两个胚珠，上胚珠败育，下胚珠可育。下胚珠倒生，具单珠被，厚珠心。大孢子母细胞减数分裂形成线性排列的四个大孢子，雌配子体发育属蓼型。开花当天，花粉散开，雌配子体尚未成熟，处

STUDIES ON THE MEGASPOROGENESIS AND MICROSPOROGENESIS AND THE DEVELOPMENT OF THEIR FEMALE AND MALE GAMETOPHYTE IN ELEUTHEROCOCCUS BRACHYPUS

The flower of Eleutherococcus brachypus has five anthers,each anther comprises four microsporangia. In meiosis of microsporocyte,cytokinesis is simultaneous type,which produces a isobilateral tetrad. Anther wall includes epidermis,endothecium, middle layer and tapetum,which belongs to dicotyledonous type. Tapetum with binucleate cells is of the secretory type. The mature pollen grain is tricelluar. The ovary is five-loculus with two ovules in each. The upper ovule is sterile and the lower ovule is fertile. The latter is mono-integinous, crassinucellate and anotropus. The megasporocyte undergoes meiosis and give rises to linear tetrad of megaspore. The development of female gametophyte is of Polygonum type. The first day in bloom, pollen grains shed,female gametophyte is not mature, which is in the four-celled embryo sac stage. E. brachypus is a protandrous plant. The relation between the development of stamen and pistil was documented.