Affiliation: | 1. Laboratory of Animal Ecology, Department of Biology , Joetsu University of Education , 1 Yamayashiki‐machi, Joetsu‐shi, Niigata 943‐8512, Japan gutturalis.39@gmail.com;3. Graduate School of Life and Environmental Sciences , University of Tsukuba , Tsukuba, 1‐1‐1 Tennoudai, Tsukuba‐shi, Ibaraki 305‐8572, Japan;4. Laboratory of Animal Ecology, Department of Biology , Joetsu University of Education , 1 Yamayashiki‐machi, Joetsu‐shi, Niigata 943‐8512, Japan |