Affiliation: | 1. The Graduate University for Advanced Studies, 10–3 Midoricho, Tachikawa, Tokyo 190–8518, Japan E‐mail: tiwata@nipr.ac.jp;2. Graduate School of Veterinary Medicine, Hokkaido University, N18W9 North Ward, Sapporo 060–0818, Japan;3. National Institute of Polar Research, 10–3 Midoricho, Tachikawa, Tokyo 190–8518, Japan and The Graduate University for Advanced Studies, 10–3 Midoricho, Tachikawa, Tokyo 190–8518, Japan;4. British Antarctic Survey, Natural Environment Research Council, High Cross, Madingley Road, Cambridge CB3 0ET, United Kingdom;5. National Institute of Polar Research, 10–3 Midoricho, Tachikawa, Tokyo 190–8518, Japan |