Affiliation: | 1. The Graduate University for Advanced Studies,10–3 Midoricho,Tachikawa, Tokyo 190–8518, JapanE‐mail: tiwata@nipr.ac.jp;2. Graduate School of Veterinary Medicine,Hokkaido University,N18W9 North Ward,Sapporo 060–0818, Japan;3. National Institute of Polar Research,10–3 Midoricho,Tachikawa, Tokyo 190–8518, JapanandThe Graduate University for Advanced Studies,10–3 Midoricho,Tachikawa, Tokyo 190–8518, Japan;4. British Antarctic Survey,Natural Environment Research Council,High Cross, Madingley Road,Cambridge CB3 0ET, United Kingdom;5. National Institute of Polar Research,10–3 Midoricho,Tachikawa, Tokyo 190–8518, Japan |