Martiodrilus olivaceous and M. panamensis, new earthworms from Colombia and Panama (Oligochaeta, Glossoscolecidae)

Two large (500-700 mm) species of Martiodrilus Michaelsen, 1937 are described from central Panama and southern Colombia. Both have polystomate nephridia, some reduced head segments, and complex folded typhlosoles. The first two conditions arc known from M. crassus Rosa 1896 and Thamnodriloides yunkeri Gates 1968. The systematic importance of polystomate nephridia and segment loss cannot be resolved without further information. Segment loss is common and variable within other glossoscoleeid genera. The two species differ as follows, the characteristics of M. olivaceous given first: tubercula pubertatis in xvii-xxv vs xviii-xxiv; small blood vessels parallel to the commissures and connected to the calciferous glands in vii-ix vs lacking these vessels; dorsal trunk ending in v vs ending in vii; intestinal nephridia with 40-50 nephrostomes vs 20-30; peptonephridia all stomate vs peptonephridia astomate in iv-vii; genital setae with 11 gouges in each of three rows, gouges ending 75 microns from tip vs 9 or 10 gouges per row, ending 200 microns from tip; spermathecae 1-4 pairs per segment each with its own duct vs 2 pairs per segment, the two of a side sharing a common duct.