State Key Laboratory Breeding Base for Zhejiang Sustainable Pest and Disease Control, Zhejiang Academy of Agricultural Sciences, Hangzhou, China;2. State Key Laboratory of Rice Biology & Ministry of Agriculturaland Rural Affairs, Key Laboratory of Molecular Biology of Crop Pathogens and Insects, Institute of Insect Sciences, Zhejiang University, Hangzhou, China;3. State Key Laboratory of Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Academy of Sciences, Beijing, China;4. Division of Plant Sciences, College of Agriculture, Food and Natural Resources, University of Missouri, Columbia, Missouri, USA;5. USDA Agricultural Research Service, Biological Control of Insects Research Laboratory, 1503 S. Providence Rd, Columbia, Missouri, USA