Review and revision of the West Timor Permian Graphiocrinus species of Johannes Wanner

In a series of papers published between 1916 and 1949 Johannes Wanner assigned or reassigned more than 1700 crinoid specimens to 26 species or subspecies of Graphiocrinus. Nine of the 26 species were subsequently transfered to other cladid genera by Wanner or other authors. Study of the amalgamated Netherlands collections of the West Timor crinoids in Naturalis Biodiversity Center in Leiden, resulted in recognition that few of the West Timor Graphiocrinus species unquestionably belonged to the genus. Review of these species and subspecies resulted in reassignment of 12 species to other genera, whereas other species and individual specimens are considered indeterminate members of several families because they lack the arms preventing positive generic identifications. This study also resulted in significant revision of the Erisocrinoidea.New taxa introduced are: Graphiocrinoidea n. superfam., Rautscholdticrinus n. gen., and Ekmelocrinus n. gen. Reassigned taxa are: Rautscholdticrinus indicus (Wanner, 1916) n. comb.; Rautscholdticrinus weidnerii (Wanner, 1937) n. comb.; Graffhamicrinus? crassus (Wanner, 1924) n. comb.; Neocatacrinus? depressus (Wanner, 1916) n. comb.; Ekmelocrinus amplior (Wanner, 1924) n. comb.; Ekmelocrinus subamplior (Wanner, 1949) n. comb.; Ekmelocrinus ovoides (Wanner, 1949) n. comb.; Ekmelocrinus verbeeki (Wanner, 1916) n. comb.; Ekmelocrinus vermistriatus (Wanner, 1916) n. comb., nomen correctum; Permiocrinus pumilus (Wanner, 1916) n. comb.; Permiocrinus quinquelobus (Wanner, 1916) n. comb.; and Apographiocrinus? rugosus (Wanner, 1916) n. comb.