Affiliation: | School of the Environment and Natural Resources, Bangor University, Bangor LL57 2UW, UK;;ESSA-Forêts, Universitéd'Antananarivo, BP 175, Antananarivo 101, Madagascar;and;Division of Biology, Imperial College London, Silwood Park Campus, Manor House, Buckhurst Road, Ascot SL5 7PY, UK |